Gilliesieae
| Gilliesieae | |
|---|---|
| |
| Ipheion Uniflorum | |
Научная классификация 
| |
| Королевство: | Plantae |
| Клада : | Трахеофиты |
| Клада : | Покрытосеменные |
| Клада : | Монокоты |
| Заказ: | Аспарагал |
| Семья: | Amaryllidaceae |
| Subfamily: | Allioideae |
| Tribe: | Gilliesieae Baker, J. Linn. Soc. London, Bot. 14: 509, 1875 |
| Type genus | |
| Gilliesia Lindl.
| |
| Genera | |
|
See text | |
| Synonyms | |
| |
Gilliesieae - это племя травянистых растений , геофитных принадлежащих к подсемейству Allioideae Амариллис семейства ( Amaryllidaceae ). Описанный в 1826 году, он содержит пятнадцать родов и около восьмидесяти видов. Это по -разному рассматривалось как подсемейство или племя. Он родом из южной части Соединенных Штатов, Центральной и Южной Америки, преимущественно Чили . Из трех племен родов, которые составляют подсемейство Allioideae, Gilliesieae является крупнейшей и наиболее переменной. Племя было разделено на два племена в 2014 году, Gilliesiae SS и Leucocoryneae, основанные на различиях в цветочной симметрии и нектарах перегородки .
Описание
[ редактировать ]Gilliesieae are perennial herbaceous geophytes characterised by simple or prolific bulbs, sometimes with lateral rhizomes. Leaf sheaths are long, tepals are more or less fused and the corona is absent. Spathe formed from 1–2 bracts. The style is more or less gynobasic. The ovary usually has two ovules per locule, side by side. There are 2–3 stamens. The commonest chromosome number is x=4. Gilliesiae is distinguished from Leucocoryneae by zygomorphic floral symmetry and the absence of septal nectaries. By contrast Leucocoryneae are zygomorphic and have septal nectaries.[1] [2]
Leucocorynae
[edit]Leucocoryneae are terrestrial perennial herbaceous plants. They have tunicate bulbs, which may be simple or prolific (with bulbils), rarely lateral rhizomes. The outer bulb scales (cataphyll) are papyraceous, colourless or violaceous (Zoellnerallium). They may or may not have a garlic like odour. The leaves are large, with membranous sheaths, usually forming an underground neck. The leaf lamina is flat, green, and glaucous, glabrous or papillose. The inflorescence may be pauciflor (Ipheion, Beauverdia, rarely Tristagma) or pluriflor (up to 30). The spathe is formed by a single bifid membranous bract (Ipheion) or from two papyraceous bracts partially fused at the base. The pedicels, which are not articulated at the receptacles, are papilose or glabrous. The flowers are hermaphroditic and actinomorphic, the perianth corolla like, with 6 (8 in Beauverdia) tepals fused at their base to form a floral tube arising around the ovary. There are 6 stamens (8 in Beauverdia), 3 fertile and 3 not (staminodes), rarely 6 (Leucocoryne), in two whorls of three (Tristagma, Ipheion) or one whorl. The filaments which are adnate (fused) to the tepals, uniting at their bases, the anthers dorsifixed (attached at their back) are oblong, yellow brown or green. The ovaries are superior and sessile with three (four in Beauverdia) carpels and locules (four in Beauverdia) and septal nectaries. The number of ovules is either 2, 4 or 30 per locule, arranged in two rows. The style is apical and persistent. The stigma has three (four in Beauverdia) lobes, or is trifid, and is papillose. The capsule, which is humifuse (Ipheion, Beauverdia) or aerocarpic, globose or prismatic, and contains many seeds (pluriseeded) which are irregular and polyhedral with a black tegmen. The embryo is linear or slightly curved.[3][4][2]
Taxonomy
[edit]
History
[edit]Lindley described Gilliesia in 1826, after fellow botanist John Gillies, placing this genus and another Chilean genus Miersia which he described at the same time, in a new taxon, Gilliesieae.[5][6][7] These and related genera have been variously assigned to families Liliaceae, Amaryllidaceae, Alliaceae and even Gilliesiaceae over their history, often as tribe Gilliesieae Lindl.[7][8] In 1985, Dahlgren's treatment of the Alliaceae (now Allioideae) within the monocotyledons, recognised three subfamilies; Gilliesioideae (Lindl.) Am., together with Agapanthoideae and Allioideae.[9] These corresponded to Hutchinson's (1959) three tribes within his expanded Amaryllidaceae (Agapantheae, Allieae, and Gilesieae).[10] The Gilliesioideae contained nine genera endemic to the southern part of South America, predominantly Chile.
Phylogenetic era (subfamily Gilliesioideae) 1996
[edit]In 1996, a molecular phylogenetic study of the rbcL gene showed Gilliesia and related genera clustering in a separate clade at subfamilial level.[8] The authors rejected the proposal of Traub (1982)[11] of a separate family, Gilliesiaceae (later resurrected by Ravenna[12]), but rather created the Gilliesioideae, as one of three subfamilies within Alliaceae, together with Allioideae and Tulbaghioideae. As phylogenetically constructed, Gilliesioideae (Gilliesioideae (Lindl.) Am., Botany: 134. 1832 - Gilliesieae Lindl. in Bot. Reg.: ad t. 992. 1826. - Type: Gilliesia Lindl.) consisted of those New World Alliaceae not included in the other two subfamilies, which included both the former Gilliesieae (Ancrumia, Erinna, Gethyum, Gilliesia, Miersia, Solaria and Trichlora) together with Ipheion, Leucocoryne, Nothoscordum, and Tristagma. Garaventia and Steinmannia were not included in the study, but considered to be part of this newly reconstructed subfamily, a total of 13 genera. [8] This is the circumscription which the Angiosperm Phylogeny Group (APG) accepted in the APG classification of 1998 and which later became known as Alliaceae sensu stricto (s.s.).[13] In the 2003 update (APGII) it was proposed to include Agapanthaceae and Amaryllidaceae under Alliaceae, while recognising an argument for renaming the overarching family from Alliaceae to Amaryllidaceae.[14]
This construction of Gilliesioideae, which represented nearly all the Alliaceae genera (i.e. except Allium and Tulbaghia), implicitly recognised that it was composed of two groups or tribes, informally referred to as Ipheieae and Gilliesieae. The Ipheieae were actinomorphic, and included Ipheion, Nothoscordum, Leucocoryne s.l. (including Pabellonia and Stemmatium). Gilliesieae were rare, mostly zygomorphic, mostly endemic to Chile and typified by Gilliesa. It contained about eight genera (Ancrumia, Gethyum, Gilliesia, Miersia, Schickendantziella, Solaria and Trichlora).[15] The genera of Gilliesioideae were thus morphologically and genetically diverse, which has made generic delimitation problematic and many species have at times been included in various different genera, and a number of genera have been shown to be polyphyletic. Consequently, the number of genera included tends to be variable.[15][7]
A more detailed analysis using multiple markers (Fay et al. 2006) confirmed the monophyly of Gilliesioideae as a whole, as were the two tribes, although some genera such as Ipheion and Nothoscordum were biphyletic.[15] In general the Gilliesieae, with their unusual floral morphology, have genera that are closely related. For instance Ancrumia, Gethyum and Solaria have been treated as three, two or one (Solaria) genus by different authors[16][1][15] (see Genera and notes).
APG III familial realignment (tribe Gilliesieae) 2009
[edit]In 2009, Chase et al. more formally brought together the three families, Agapanthaceae, Alliaceae, Amaryllidaceae, under the single Asparagalean monophyletic family, now renamed Amaryllidaceae from Alliaceae, reversing the Dahlgrenian process of family splitting. This necessitated reducing the existing ranks of the component subfamilial taxa. [17] This formed the basis for the 2009 APG classification (APGIII).[18] Thus subfamily Gilliesioideae became tribe Gilliesieae (Baker, J. Linn. Soc., Bot. 14: 509. 24 Apr 1875) within subfamily Allioideae of family Amaryllidaceae. Within the tribe they included thirteen genera including Leucocoryne s.l. (see Genera).
The full taxonomy of tribe Gilliesieae remains unresolved. Of the South America genera, a number have common features (tunicate bulbs, inflorescences with unarticulated pedicels, and one or two bracts subtending the inflorescence). These are Ipheion Raf., Leucocoryne Lindl., Nothoscordum Kunth, Tristagma Poepp., and Zoellnerallium Crosa. The position of Ipheion is particularly problematic.[4]
Division of Ipheion (resurrection of Beauverdia)
[edit]In 1972, Ipheion was divided into two sections, Hirtellum and Ipheion. However, the development of phylogenetic analysis revealed that Ipheion was not monophyletic, although the division into sections was later supported. Beauverdia Herter had been first described in 1943.[19] Originally, it was created to distinguish those species with unifloral inflorescences from others with plurifloral inflorescences within Nothoscordum and other genera, no longer considered Amaryllidaceae. As proposed, it had ten species but its independence was short lived, being returned to a synonym of Ipheion, and a number of species were transferred to other genera, including Nothoscordum and Tristagma.[4]
In 2014, Ipheion section Hirtellum was again raised to genus rank and restored to the tribe, being distinguished from other Ipheion species, under the older name of Beauverdia, with four species found in Argentina, southern Brazil, and Uruguay.[4]
Division of Gilliesieae and resurrection of Leucocoryneae
[edit]In 2014 Sassone also proposed resurrecting an older taxon, Leucocorynae to include six genera, Beauverdia (4 species), Ipheion s.s. (3 species), Leucocoryne s.l. (15 species), Nothoscordum (c. 20 species), Tristagma (c. 20 species) and Zoellnerallium (2 species).[2] Leucocorynae had originally been described by Ravenna in 2001 as a tribe of Gilliesioideae, to include Leucocoryne together with Tulbaghia (now in separate tribe, Tulbaghieae) on morphological grounds, but it was not adopted. Instead, as described by Rudall et al. (2002)[16] and Fay et al. (2006)[15] there was a general recognition, as described above of two tribes, Ipheieae nom. nud. (4 genera) and Gilliesieae (7 genera) differing by actinomorphic floral symmetry and the presence of septal nectaries in the former.[16][2] Subsequently, Zoellnerallium was added to the Ipheieae,[20][21] even though with the reduction of Gilliesioideae to the tribe Gilliesieae, the older divisions could no longer be recognised, at least as tribes (possibly subtribes).[22]
This now formally divides the tribe Gilliesieae s.l. into two tribes, Gilliesieae s.s. (8 genera) and Leucocoryneae (6 genera). This new tribe corresponds to the older Ipheieae, together with the two more recent additions of Beauverdia and Zoellnerallium and includes about 65 species, although this could be closer to 130, according to Ravenna's proposals for Nothoscordum which would increase its species from 20 to about 60.[2][23]
The taxonomy of Gilliesieae s.s. remains difficult with limited sampling, because of the problem of obtaining material from these little-known plants. Hence the different treatment of a number of the genera by different authors.[15] (see Genera and notes)
Genera
[edit]Included genera
[edit]Included genera according to Chase et al., [17] as modified by Sassone et al. 2014[4][2] and García et al. 2022.[24]
- Tribe Leucocoryneae (Ipheieae group) (Ravenna) Sassone, S.C. Arroyo & Giussani[25]
- Atacamallium Nic.García (2022)[24]
- Beauverdia Herter (1943).[4]
- Ipheion Raf. (1836).
- Leucocoryne Lindl. (1830).
- Nothoscordum Kunth (1843).
- Tristagma Poepp. (1833).
- Zoellnerallium Crosa (1975).
- Tribe Gilliesieae s.s.
- Ancrumia Harv. ex Baker (1877).[notes 1]
- Erinna Phil. (1864).[notes 2]
- Gethyum Phil. (1873).[notes 3]
- Gilliesia Lindl. (1826). Type species
- (including Pabellonia Quezada & Martic. and Stemmatium Phil.)
- Miersia Lindl. (1826).
- Schickendantziella Speg. (1903).
- Solaria Phil. (1858).
- Speea Loes. (1927).
- Trichlora Baker (1877).
Uncertain, doubtful or former genera
[edit]Three genera have been transferred to Allium.[26] Caloscordum Herb. (1844)., which is now more properly considered part of Allium,[15] Both Herbert (1844)[27] and Lindley (1847)[28] had originally considered it a distinct genus, while others considered it as part of Nothoscordum.(Li 1996)[29][15] Milula is embedded in Allium as a section.[15][30] Garaventia is considered part of Tristagma.[31] Muilla was included in the Allioideae by Dahlgren,[32] but in tribe Brodiaeeae. That tribe was subsequently raised to family status as Themidaceae.[8]
- Caloscordum Herb. (1844). (Subgenus of Allium)[33]
- Garaventia Looser (1941). (syn. Tristagma)
- Milula Prain (1896). (Section of Allium)
- Muilla S.Watson ex Bentham (1883). (Themidaceae)
- Nectaroscordum Lindl. (1836). (Subgenus of Allium)[33]
Species
[edit]There are about eighty species included in the tribe.[2]
Distribution
[edit]The Gilliesieae are endemic to the southern part of South America, predominantly Chile.[8] The Leucocoryneae are also a South American tribe with the exception of two species of Nothoscordum (N bivalve, N. gracile) which extend to southern North America, otherwise they are found in southern Brazil, Argentina, Uruguay and Chile.[2] (see map in Stevens 2013).[34]
Notes
[edit]- ^ Ancrumia: Rahn (1998) considered Ancrunia to be part of Solaria (Kubitzki 1998, Rahn: Alliaceae. pp. 70–78) but both (Zöllner & Arriagada 1998) and (Rudall et al. 2002) considered them as separate genera. As of 2014, the World Checklist considers the single species of this genus to be a synonym of Solaria cuspidata, and does not accept Ancrumia as a separate genus (see discussion above). (WCSP 2015, WCSP Ancrumia)
- ^ Erinna: Described by Philippi in 1864 as a monotypic genus, based on Erinna gilliesioides. (Philippi 1864, 1073. Erinna. p. 266) As such it was a genus within Alliaceae, and included in the phylogenetic construction of Gilliesieae in 1996.(Fay & Chase 1996) Although (Ravenna 2000) proposed transferring it to Leucocoryne on morphological grounds, (Sassone et al. 2014b) it was included separately by (Chase et al. 2009) and hence the 2009 APGIII. (APG 2009) (Sassone et al. 2014b) Although the World Checklist lists Erinna as a synonym of Leucocoryne, (WCSP 2015, Erinna Phil.) (Sassone et al. 2014b) still considered its status uncertain.
- ^ Gethyum: Rahn (1998) considered Gethyum to be part of Solaria, (Kubitzki 1998, Rahn: Alliaceae. pp. 70–78) but both (Zöllner & Arriagada 1998) and (Rudall et al. 2002) considered them as separate genera and Gethyum was included in the 2009 construction of Gilliesiea, (Chase et al. 2009) as discussed above. However the World Checklist considers it part of Solaria. (WCSP 2015, Gethyium)(Fay & Hall 2007)
References
[edit]- ^ Jump up to: a b Fay & Hall 2007.
- ^ Jump up to: a b c d e f g h Sassone et al. 2014b.
- ^ Sassone et al. 2013.
- ^ Jump up to: a b c d e f Sassone et al. 2014a.
- ^ Lindley 1826.
- ^ Lindley 1846, CCXLVIII Gilliesieae. pp. 275-277.
- ^ Jump up to: a b c Rix et al. 2013.
- ^ Jump up to: a b c d e Fay & Chase 1996.
- ^ Dahlgren, Clifford & Yeo 1985.
- ^ Hutchinson 1959.
- ^ Traub 1982.
- ^ Ravenna 2000a.
- ^ APG 1998.
- ^ APG 2003.
- ^ Jump up to: a b c d e f g h i Fay, Rudall & Chase 2006.
- ^ Jump up to: a b c Rudall et al. 2002.
- ^ Jump up to: a b Chase et al. 2009.
- ^ APG 2009.
- ^ Herter 1943.
- ^ Crosa 1975.
- ^ Crosa 2004.
- ^ Escobar et al. 2012.
- ^ The Plant List 2013, Nothoscordum.
- ^ Jump up to: a b García, N., Sassone, A.B., Pinto, R. and Román, M.J. (2022), Atacamallium minutiflorum (Amaryllidaceae, Allioideae), new genus and species from the coastal desert of northern Chile. TAXON, 71: 552-562. https://doi.org/10.1002/tax.12684
- ^ Ravenna 2001.
- ^ Friesen, Fritsch & Blattner 2006.
- ^ Herbert 1844.
- ^ Lindley 1847.
- ^ Li et al. 1996.
- ^ WCSP 2015, Milula.
- ^ WCSP 2015, Garaventia.
- ^ Dahlgren, Clifford & Yeo 1985, Brodiaeeae p. 196 .
- ^ Jump up to: a b Li et al. 2010.
- ^ Stevens 2013.
Bibliography
[edit]General
[edit]- Lindley, John (1846). An Introduction to the Natural System of Botany: or, A Systematic View of the Organisation, Natural Affinities, and Geographical Distribution, of the Whole Vegetable Kingdom. London: Bradbury. Retrieved 26 January 2015.
- Philippi, RA (1864). "Plantarum novarum Chilensium, inclusis quibusdam Mendocinis et Patagonicis". Linnaea. 33: 1–308. Retrieved 21 January 2015.
- Hutchinson, John (1959). The families of flowering plants, arranged according to a new system based on their probable phylogeny. 2 vols. Macmillan.
- Dahlgren, R.M.; Clifford, H.T.; Yeo, P.F. (1985). The families of the monocotyledons. Berlin: Springer-Verlag. ISBN 978-3-642-64903-5. Retrieved 10 February 2014. Available on Google Books
- Kubitzki, K., ed. (1998). The families and genera of vascular plants. Vol.3. Berlin, Germany: Springer-Verlag. ISBN 978-3-540-64060-8. Retrieved 14 January 2014.
- Stevens, P.F. (2013), "Asparagales: Amaryllidaceae", Angiosperm Phylogeny Website
- The Angiosperm Phylogeny Group (1998). "An ordinal classification for the families of flowering plants" (PDF). Annals of the Missouri Botanical Garden. 85 (4): 531–553. doi:10.2307/2992015. JSTOR 2992015. S2CID 82134384. Archived from the original (PDF) on 2011-06-08.
- The Angiosperm Phylogeny Group (April 2003). "An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants: APG II". Botanical Journal of the Linnean Society. 141 (4): 399–436. doi:10.1046/j.1095-8339.2003.t01-1-00158.x.
- Angiosperm Phylogeny Group (2009), "An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants: APG III", Botanical Journal of the Linnean Society, 161 (2): 105–121, doi:10.1111/j.1095-8339.2009.00996.x, hdl:10654/18083
- Chase, Mark W.; Reveal, James L.; Fay, Michael F. (October 2009). "A subfamilial classification for the expanded asparagalean families Amaryllidaceae, Asparagaceae and Xanthorrhoeaceae". Botanical Journal of the Linnean Society. 161 (2): 132–136. doi:10.1111/j.1095-8339.2009.00999.x.
- Rina Kamenetsky; Hiroshi Okubo, eds. (2012). Ornamental Geophytes: From Basic Science to Sustainable Production. CRC Press. ISBN 978-1-4398-4924-8.
- Howard, Thad M. (2001). Bulbs: From Warm Climates. Austin: University of Texas. ISBN 978-0292731264. Retrieved 23 January 2015.
Amaryllidaceae (Gilliesieae)
[edit]- Lindley, John (1826). "Gilliesia graminea". Edwards' Botanical Register. 12: t. 992. Retrieved 19 January 2015.
- Hutchinson, J (1939). "The tribe Gilliesieae of Amaryllidaceae". Herbertia. 6: 136–145.
- Huber, Herbert F.J. (1969). "Die Samenmerkmale und Verwandtschaftsverhältnisse der Liliiflorae". Mitt. Bot. Staatssamml. [Mitteilungen der Botanischen Staatssammlung München]. 8: 219–538. Retrieved 10 February 2015.
- Traub, H. P. (1982). "Order Alliales". Plant Life. 38: 119–132.
- Fay, Michael F.; Chase, Mark W. (August 1996). "Resurrection of Themidaceae for the Brodiaea alliance, and Recircumscription of Alliaceae, Amaryllidaceae and Agapanthoideae". Taxon. 45 (3): 441–451. doi:10.2307/1224136. JSTOR 1224136.
- Fay, MF; Rudall, PJ; Chase, MW (2006). "Molecular Studies of Subfamily Gilliesioideae (Alliaceae)". Aliso. 22: 367–371. doi:10.5642/aliso.20062201.30. Retrieved 7 January 2015.
- Escobar, Inelia; Ruiz, Eduardo; Baeza, Carlos (2012). "Estudios cariotípicos en especies de Gilliesieae Lindl. (Gilliesioideae-Alliaceae) de Chile central". Gayana. Botánica. 69 (2): 240–250. doi:10.4067/S0717-66432012000200003.
- Dutilh, Jha (2009). «Нетропические Alliaceae» . Milliken, W., Klitgård, B. & Baracat, A. neotropikey - Интерактивный ключ и информационные ресурсы для цветущих растений неотропиков . Получено 8 января 2015 года .
- Ли, Q.-Q.; Zhou, S.D.; Он, X.-J.; Yu, Y.; Zhang, Y.-C.; Вей, X.-Q. (21 октября 2010 г.). «Филогения и биогеография Allium (Amaryllidaceae: Allieae) на основе ядерной рибосомной внутренней транскрибированной проставки и последовательностей хлоропластов RPS16, фокусируясь на включении видов, эндемичных для Китая» . Анналы ботаники . 106 (5): 709–733. doi : 10.1093/aob/mcq177 . PMC 2958792 . PMID 20966186 .
- Рудалл, PJ; Бейтман, RM; Fay, MF; Истман А. (1 декабря 2002 г.). «Цветочная анатомия и систематика Alliaceae с особой ссылкой на Гиллисию , предполагаемое насекомое, имитирующее с сильно зигоморфными цветами». Американский журнал ботаники . 89 (12): 1867–1883. doi : 10.3732/ajb.89.12.1867 . PMID 21665616 .
- Фризен, N (ноябрь 2000 г.). «Молекулярные и морфологические доказательства происхождения аберрантного рода Милулы в гималайских видах Allium (Alliacae)». Молекулярная филогенетика и эволюция . 17 (2): 209–218. doi : 10.1006/mpev.2000.0844 . PMID 11083935 . S2CID 27171819 .
- Фризен, Николай; Фрич, Рейнхард М.; Блаттнер, Фрэнк Р. (2006). «Филогения и новая внутригенерическая классификация allium (alliaceae) на основе ядерной рибосомальной ДНК ее последовательностей» . Алисо . 22 : 372–395. doi : 10.5642/aliso.20062201.31 . Получено 8 января 2015 года .
- Сассоне, Агостина Б.; Аррой-Леюенбергер, Сильвия С.; Джуссани, Лилиана М. (31 декабря 2014b). «Nueva incripción de la tribu Leucocoryneae (amaryllidaceae, allioideae)» . Дарвиниана . Nueva Series. 2 (2): 197–206. Doi : 10.14522/darwiniana/2014.22.584 . ISSN 0011-6793 . Получено 9 Janogy 2015 .
- Торрес-Меллада, Густаво А; Эскобар, Инелия; Пэлфнер, Гоц; Казанова-Кань, М. Анжелика (июнь 2012 г.). «Микотрофия в Gilliesieae, угрожаемое и плохо известное племя Alliaceae из центрального Чили» . Чилийский журнал естественной истории . 85 (2): 179–186. Doi : 10.4067/s0716-078x2012000200004 .
- Zöllner, O.; Arriada, L. (1998). Племя Gilliesiae находится в Чили. Гербертия 53 : 104–107.
- Равенна, с. (2000a). «Семейство Gilliesiaceae». Онира ботанических листовок . 4 : 11–14.
Роды
[ редактировать ]- Хертер, WG (1943). « Бовердия вроде нового Liliacearum». Боисьера . 7 : 505-512.
- Герберт, W (1844). "64. Caloscordum" . Ботанический регистр Эдвардса . 30 : 66–70 . Получено 20 января 2015 года .
- Линдли, Дж. (1847). «Caloscordum Nerinefolium. Nerine-Leared Caloscord» . Ботанический регистр Эдвардса . 33 : Тарелка 5 . Получено 20 января 2015 года .
- Li, RJ; Шан, Зи; Cui, TC; JM Xu., JM (1996). «Исследования по кариотипам и филогенетической взаимосвязи секты Allium . Caloscordum (Liliaceae) из Китая» . Acta Phytotax. Грех (на китайском). 34 (3): Acta Phytotax. Грех
- Фэй, Майкл Ф.; Холл, Тони (май 2007). «589. Гетюм Atropurpureum». Кертис от ботанического журнала . 24 (2): 121–126. doi : 10.1111/j.1467-8748.2007.00573.x .
- Рикс, Мартин; Странно, комплект; Рудалл, Паула Дж. (Апрель 2013 г.). «753. Гиллисия Монтана». Кертис от ботанического журнала . 30 (1): 28–35. doi : 10.1111/curt.12014 .
- Сассоне, Агостина Б.; Джуссани, Лилиана М.; Гуаглианон, Энкорносион Р. (30 апреля 2013 г.). «Многофакторные исследования Ipheion (Amaryllidaceae, Allioideae) и связанные с ними генерируются». Систематика растения и эволюция . 299 (8): 1561–1575. Doi : 10.1007/s00606-0819-5 . HDL : 11336/18970 . S2CID 254058196 .
- Сассоне, Агостина Б.; Джуссани, Лилиана М.; Гуаглианон, Энкорн Ан Р. (1 июля 2014a). « Бовердия , всплывший род amaryllidaceae (allioideae, gilliesieae)». Систематическая ботаника . 39 (3): 767–775. Doi : 10.1600/03636414x681527 . HDL : 11336/19039 . S2CID 86287897 .
- Равенна, П.Ф. (2000). «Новые или достойные внимания виды Leucocoryne (Alliaceae)». Онира . 4 : 3–10.
- Равенна, ПФ (2001). «Новые или достойные внимания виды Leucocoryne (Alliaceae) III». Онира . 5 : 42–43.
- Jara-Arancio, Paola; Арройо, Мэри Тк; Герреро, Пол С.; Hinojosa, Luis F.; Арансио, Джина; Мендес, Марко А.; Карин, Марк (февраль 2014 г.). «Филогенетические перспективы на сдвиги биомема в лейкокорине (Alliaceae) в отношении эволюции климатической ниши в Западной Южной Америке» (PDF ) Журнал биогеографии 41 (2): 328–3 Doi : 10.1111/jbi.12186 . S2CID S2CID Получено 9 января
- Crosa, или (1975). « Zoellnerallium , Género Nuevo Para La Tribu Allieae (Liliaceae)». Дарвиниана (на испанском). 19 : 331–334.
- Crosa, O (2004). «Секулональные виды и гендерные гендерные zoelelliut (alliaceae) » Дарвиниан (на испанском). 42 (1–4): 165–1 Получено 20 января
Базы данных
[ редактировать ]- "Дом" . Список заводов (2013). Версия 1.1 . 2013 . Получено 4 января 2015 года .
- "Быстрый поиск" . Всемирный контрольный список избранных семейных семей . Королевские ботанические сады, Кью . Получено 8 января 2015 года .
 Подразделения семейства Amaryllidaceae с родом типа |
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