Schaly Brrasted Munia
| Schaly Brrasted Munia | |
|---|---|
| |
| Л. П. Punctulata ( Индия ) | |
Научная классификация 
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| Домен: | Эукариота |
| Королевство: | Животное |
| Филум: | Chordata |
| Сорт: | Птица |
| Заказ: | Прохожий |
| Семья: | Estrildidae |
| Род: | Lonchura |
| Разновидность: | L. Point
|
| Биномиальное название | |
| Пунктурный Lonchura | |
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| Местный диапазон | |
| Синонимы | |
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Шешенистая муния или пятнистая муния ( Lonchura Punctulata ), известная в торговле домашних животных как мускатный орех Mannikin или Spice Finch , является воробьем , родом из эстральдидного зяблика в тропической Азии. Вид рода Lonchura , он был официально описан и назван Карлом Линнеусом в 1758 году. Его название основано на различных масштабных маркировках перьев на груди и животе. Взрослый коричневый выше и имеет темный конический счет. Вид имеет 11 подвидов по всему диапазону, которые немного различаются по размеру и цвету.
Эта муния ест в основном семена травы, кроме ягод и небольших насекомых. Они добывают в стаях и общаются с мягкими звонками и свистками. Вид очень социальный и может иногда укорениться с другими видами мунией. Этот вид встречается на тропических равнинах и лугах . Перецены пары конструируют купольные гнезда, используя траву или бамбуковые листья.
Этот вид эндемичен по отношению к Азии и происходит от Индии и Шри -Ланки на восток до Индонезии и Филиппин (где он называется Mayang Pakíng ). Он был введен во многие другие части мира, и дикие группы населения зарекомендовали себя в Пуэрто -Рико и Хиспаньоле , а также в частях Австралии и Соединенных Штатов Америки с наблюдениями в Калифорнии. Птица перечисляется как наименьшая забота Международным союзом сохранения природы (МСОП).
Таксономия
[ редактировать ]В 1743 году английский натуралист Джордж Эдвардс включил иллюстрацию и описание чешуйчатой мунии в первом томе своей естественной истории необычных птиц . Он использовал английское имя «Говри птица». Эдвардс основал свое офорта в ручном цвете на образце в лондонском доме Чарльза дю Буа , казначея в Ост-Индской компании . [ 2 ] Когда в 1758 году шведский натуралист Карл Линнеус обновил свою систему Naturae для десятого издания , он поместил заглашенную мунию с перекрестными путями в роду Loxia . Linnaeus включил краткое описание, придумал биномиальное название Loxia punctulata и процитировала работу Эдвардса. [ 3 ] Линнеус указал место как «Азию», но это было ограничено Калькуттой (Калькутта) Е.С. Стюартом Бейкером в 1926 году. [ 4 ] [ 5 ] В настоящее время этот вид помещается в род Lonchura , который был представлен английским натуралистом Уильямом Генри Сайксом в 1832 году. [ 6 ] [ 7 ] Название рода Lonchura сочетает в себе древнегреческий лонххи, означающий «копья-головы» или «копья» с Oura, означающим «хвост». Конкретный эпитет из современного латинского пунктулатуса, означающего «пятнистый» или «пунктирный». [ 8 ]
В большом диапазоне существует 11 признанных подвидов . К ним относятся форма назначения , обнаруженную на равнинах индийского субконтинента , включая Пакистан, Индию, Иран, Непал, Бангладеш и Шри -Ланка. Название Lineoventer раньше использовалось для индийского населения. Другие популяции включают в себя субиндураты из восточных Гималаев, Юннаненсис из южного Китая, Тпель -Таиланд, Кабаниси Филиппин и фрутис Сингапура и Суматра. Популяции островов включают Низорию (Java, Bali, Lombok, Sumbawa), Particeps (Sulawesi), Baweana (остров Бавин), Sumbae (Sumba) и Blasii (Flores, Timor и Tanimbar). [ 7 ]
- Л. П. Пунктулата ( Линнеус , 1758 ) - Северный Пакистан, Индия (кроме северо -востока), Непал Терай и Шри -Ланка
- Л. П. Subundulata ( Годвин-Остин , 1874)-Бутан, Бангладеш, Северо-Восточная Индия (Ассам) и Западная Мьянма
- Л. П. Yunnanensis Parkes, 1958 - Южный Китай (юго -восточный Xizang , South Sichuan , Yunnan ) и Северная Мьянмар
- L. p. topela (R. Swinhoe, 1863) – southern Myanmar, Thailand, southeast China (Taiwan), Hainan Islands, Laos, Cambodia and Vietnam
- L. p. cabanisi (Sharpe, 1890) – north and west Philippines (Luzon, Mindoro, Calauit, Palawan, Panay, Negros, Cebu, Mindanao) and northern Borneo (coastal west Sabah and Brunei)
- L. p. fretensis (Kloss, 1931) – south Malay Peninsula, Singapore, Sumatra and Nias Islands
- L. p. nisoria (Temminck, 1830) – southern Borneo (western & southern Kalimantan), Java, Bali, and western Lesser Sundas (Lombok, Sumbawa)
- L. p. sumbae Mayr, 1944 – Sumba, in western Lesser Sundas
- L. p. blasii (Stresemann, 1912) – central & eastern Lesser Sundas (Flores east to Timor and Tanimbar Islands)
- L. p. baweana Hoogerwerf, 1963 – Bawean Islands, off northeast Java
- L. p. particeps (Riley, 1920) – Sulawesi
The subspecies holmesi (southeast Borneo) is sometimes recognised.[9]
Description
[edit]
The scaly-breasted munia is about 11–12 centimetres (4.3–4.7 in) long and weighs 12–16 grams (0.026–0.035 lb). The adult has a stubby dark bill typical of grain eating birds, brown upperparts and a dark brown head. The underparts are white with dark scale markings. The sexes are similar, although males have darker markings on the underside and a darker throat than females.[10]
Immature birds have pale brown upperparts, lack the dark head found in adults, and have uniform buff underparts that can be confused with juveniles of other munia species such as the tricolored munia (Lonchura malacca) across the Asian and island populations and the black-throated munia (Lonchura kelaarti) in parts of India or Sri Lanka.[10][11]
Distribution and habitat
[edit]
Scaly-breasted munias are found in a range of habitats but are usually close to water and grassland. In India, they are especially common in paddy fields where they are considered a minor pest on account of their feeding on grain. They are found mainly on the plains, but can be observed in the foothills of the Himalayas, in which they may be present at altitudes near 2,500 m (1.6 mi), and in the Nilgiris, where they are found at altitudes up to 2,100 m (6,900 ft) during the summer. In Pakistan, they are restricted to a narrow region from Swat in the west to Lahore, avoiding the desert zone, and then occurring again in India east of an area between Ludhiana and Mount Abu.[13] The species has also been observed in Kashmir, though this is rare.[14][15]
Outside their native range, escaped birds frequently establish themselves in areas with a suitable climate and can then colonize new areas nearby. Escaped cage-birds established in the wild and such populations have been recorded in the West Indies (Puerto Rico since 1971),[16] Hawaii (since 1883[17]),[18] Japan[19] and southern United States, mainly in Florida and California.[20][21] In Oahu, Hawaii, they compete for habitats with the tricolored munia and tend to be rare where this competitor is present.[18] The species has been introduced to other parts of the world due to its popularity as a cage bird and populations have established in the wild.[22][23]
Behaviour and ecology
[edit]Sociality
[edit]Scaly-breasted munias form flocks of as many as 100 birds. Individuals communicate with calls that include a short whistle, variations of kitty-kitty-kitty, and a sharp chipping alarm note.[11][23] They sometimes flick their tails and wings vertically or horizontally while hopping about. The tail flicking motion may have evolved from a locomotory intention movement. The exaggerated version of the tail flicking movement may have undergone ritualization. As a social signal, tail flicking in several other species acts as a signal indicating the intent to fly and helps keep flocks together.[23][24]
When roosting communally, scaly-breasted munia sit side by side in close contact with each other. The outermost bird often jostles towards the center. Birds in a flock sometimes preen each other, with the soliciting bird usually showing its chin. Allopreening is usually limited to the face and neck.[23] The scaly-breasted munia is rarely hostile but birds will sometimes quarrel without any ritualized posturing.[23]
Breeding
[edit]
The breeding season is during the summer rainy season (mainly June to August and also in October season in India) but can vary. Laboratory studies have found that long day illumination and high humidity trigger gonadal growth.[25] The song of the male is very soft but complex and variable, audible only at close range. This song described as a jingle consists of a series of high notes followed by a croaky rattle and ending in a slurred whistle. When singing the male sits in what is called the slope posture—erect with the head feathers raised.[23]
There are two types of slope posture, a pre-copulatory one and an ordinary one. The pre-copulatory behavior of scaly-breasted munia includes a sequence of actions. The first involves either the male or female playing with nest-material. As soon as a bird has arranged the nest material in its bill, it begins to fly around in a zigzag path. Once a bird lands close to its partner, the male bends towards the female and wipes its bill. The male then sings with movements of the body. The female invites mounting with tail quivering.[11][23] The nest is a large domed structure loosely woven from blades of grass, bamboo or other leaves with a side entrance and is placed in a tree or under the eaves of a house. A study in southern India found the preferred nesting trees to be Toddalia asiatica, Gymnosporia montana and Acacia chundra, especially short and bushy ones in areas with low canopy cover. The nest opening is located to face downwind of the most frequent wind direction.[26] In northern India, they preferred isolated Acacia nilotica in non-urban areas but used Thuja orientalis and Polyalthia longifolia in urban gardens.[27]
Scaly-breasted munia clutches usually contain 4 to 6 eggs, but can contain up to 10. Both sexes build the nest and incubate the eggs, which hatch in 10 to 16 days.[14][28]
The species is extensively used as a brood host by the parasitic pin-tailed whydah in Southern California — where both species are feral — with the munia raising the whydah's chicks as its own.[29] This relationship is novel, as the two species do not naturally co-occur in their native ranges, and had no established evolutionary relationship as parasite and host.[29]
Food and foraging
[edit]
The scaly-breasted munia feeds mainly on grass seeds, small berries such as those of Lantana and insects.[30] Although the bill is suited for crushing small grains, they do not show lateral movements of the lower mandible which help European greenfinches in dehusking seeds.[31] Like some other munias, they may also feed on algae, a rich protein source, prior to the breeding season.[32]
The ease of maintaining these birds in captivity has made them popular for studying behavior and physiology. Feeding behavior can be predicted by the optimal foraging theory, where animals minimize time and energy spent to maximize food intake. This theory has been tested by studying the strategies used by scaly-breasted munias to increase their feeding efficacy.[33]
Flock size tradeoffs
[edit]Studies on foraging have examined the effect of group size in reducing time spent on predator vigilance, thereby increasing feeding efficiency. According to the "many-eyes" hypothesis,[34] a reduction in the individual time spent on vigilance against threats in larger groups allows for more time to be spent on searching for food and feeding. Vigilance is greatest among solitary individuals and reduces as the group size increases to about four. The birds collect seeds more quickly in larger groups, reflecting a decrease in individual vigilance, a decrease in handling time, and an increase in both search speed and focus when foraging.[35]
Individuals may also take advantage of group foraging by "joining" members that have found food. The options to seek food or to join others that have discovered food involves information sharing and has been studied through what are termed "producer-scrounger models".[36] A cost associated with group foraging is increased resource competition, which in turn may reduce anti-predatory vigilance due to the intensity of foraging.[37] Some studies show that increased competition results in a decreased feeding rate.[38]
Foraging models
[edit]When foraging, scaly-breasted munia can search as individuals or search for others that have found food and join them. The economic consequences of the decision to join others has been modeled in two ways: the producer-scrounger model and the information sharing model. These models are based on hypotheses that differ in the degree of compatibility that is assumed between the two food and joining opportunity search modes.[39]
The information sharing model assumes that individuals search concurrently for finding and joining opportunities while the producer-scrounger model assumes that the search modes are mutually exclusive.[39] Hopping with the head facing up and downward are observed to be statistically associated with the frequencies of a bird's joining and finding, respectively. When the expected stable frequency of the scrounger tactic was altered by changing the availability of seeds, the relative frequency of hopping with the head up changed accordingly. When the seed distribution made the scrounger tactic unprofitable, the frequency of hopping with the head up diminished and appears to support the predictions of the producer-scrounger model.[40]
Studies show that scaly-breasted munias tend to adopt the scrounger tactic when food is more clumped and when the group size increases. When most foragers adopt scrounging, the time taken to discover new food patches is greater.[41]
Vigilance
[edit]Most social foragers must search for food while also avoiding predators. It has been suggested that individuals that play scrounger could also, by virtue of their head position, be alert for predators and hence contribute to antipredatory vigilance. If the scrounger tactic is compatible with antipredatory vigilance, then an increase in antipredatory vigilance should lead to the detection of more joining opportunities, and hence more joining. When stationary, the head-up tactic has been shown to be associated with antipredatory vigilance. However scanning while hopping does not aid in vigilance and it is thought that the scrounger tactic is incompatible with antipredatory vigilance in the scaly-breasted munia.[42]
Specialized foraging
[edit]Scaly-breasted munias have variable competitive behaviors that allow them to exploit scarce resources. There are two foraging alternatives: producers that make the food available and scroungers that steal food found by the producers. Studies show that these choices lead to a stable equilibrium within a group. When individuals are free to choose between producer and scrounger, frequency dependent selection results in a stable mixture of both behaviors where each receives similar payoff. Studies indicate that if most of the population consists of producers, then scrounging behavior is favored by natural selection because there is plenty of food to steal. On the other hand, if most birds exhibit scrounging then the competition for stealing is so great that producing is favored.[43][44]
Three hypotheses might account for consistent foraging specializations across individuals: food source variation, phenotypic differences, and frequency dependent-choice. The food source variation hypothesis predicts that individuals will specialize when the use of two skills is more costly than specialist foraging. The phenotypic differences hypothesis proposes that individuals differ in their ability to use each foraging skill and stably specialize on the most profitable one. The pattern of specialization is expected to be stable although the number of individuals that use a given skill depends on the phenotypic composition of the flock. The frequency dependent choice hypothesis also proposes that individuals specialize on the most profitable skill, but the profitability of each alternative decreases as the number of phenotypically identical foragers gradually specialize on each skill when initially given two equally profitable alternatives. At equilibrium, individual payoffs should be independent of the pattern of specialization. Individuals in flocks adjusted their use of the two skills and two birds in each flock specialized on a different skill resulting in a variant of both the food source variation hypothesis and frequency dependent choice hypothesis.[45]
Aviary experiments conducted with captive flocks of scaly-breasted munia have tested whether producers and scroungers reach the predicted stable equilibrium frequency (see Evolutionarily stable strategy) when individuals are free to choose either behavior. The numbers choosing either producers and scrounger strategies have been shown to converge on stable frequencies while demonstrating that variation in tactics arise through frequency dependent pay-offs from the choice of different feeding strategies.[46]
Furthermore, foraging birds may feed actively on the substrate or pick grains dropped on the ground and these strategies may be chosen according to the situation. Early departures occur more often when expected searching time decreases and when competition intensity increases. Competition intensity is expected to increase when more scroungers are present or when patches are smaller.[47]
Prey crypsis
[edit]Since producers search for food and scroungers wait for opportunities to join, prey crypsis imposes a producer specific cost that shifts the producer scrounger equilibria towards more scrounging. Prey crypsis resulted in increased latency to eat the seed and increased number of detection errors.[48] Moreover, the presence of a competitor negatively affected foraging efficiency under cryptic backgrounds. The foraging efficiency of individuals that had previously foraged with a competitor on cryptic seeds remained low even after the competitor had been removed. Thus, the costs of foraging on cryptic prey may be greater for social foragers than for solitary foragers.[49]
Resource defence
[edit]Recent models of economic defence in a group-foraging context predict that the frequency of aggressive interactions should decline as resource density increases.[50][51][52] Studies with scaly-breasted munia show that the intensity of aggressive encounters was highest when patch location was signaled, and the effect of changing resource density depended on whether patch location was signaled or not. Signaling patch location was equivalent to making the resources more spatially predictable. Changing patch density had no effect on the number of aggressive encounters when the location of food was not signaled. When food location was signaled, increasing patch density resulted in the predicted decrease in the number of aggressive encounters.[53]
Conservation
[edit]The scaly-breasted munia is an abundant species and classified as least concern on the IUCN Red List.[1] The species occupies an extremely large range, and its population, while still unquantified, is large and stable. The scaly-breasted munia is not globally threatened and is common to very common throughout most of its range. However, some populations are dwindled due to the increase of bird cagings.[54]
In many areas it is regarded as an agricultural pest, feeding in large flocks on cultivated cereals such as rice.[55] In Southeast Asia, the scaly-breasted munia is trapped in large numbers for Buddhist ceremonies, but most birds are later released.[56]
References
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