Кентисутус

Кентисутус Временный диапазон: ранний эоцен ,
К. Спенсери Ископая в Музее науки Теллуса
Научная классификация
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Clade:	Archosauromorpha
Clade:	Archosauriformes
Order:	Crocodilia
Superfamily:	Gavialoidea
Genus:	†Kentisuchus Mook, 1955
Species
†K. astrei Jouve, 2016 †K. spenceri (Buckland, 1836) [originally Crocodylus spenceri] (type)
Synonyms
Crocodylus spenceri Buckland, 1836

Кентисутус является вымершим родом гэвилоидного крокодилиана , традиционно считается членом подсемейства Tomistominae . Окаменелости были найдены из Англии и Франции , которые датируются ранним эоценом . Род также был записан из Украины , но неясно, относятся ли образцы из Украины к Кентисухусу .

Разновидность

К. Спенсери Череп

Род Kentisuchus был возведен Чарльзом Муком в 1955 году для вида «Crocodylus» Toliapicus , описанного Ричардом Оуэном в 1849 году. Уильям Бакленд назвал «Crocodylus» Спенсери на основе частичного черепа, найденного на острове Шеппи в Кент , Англия Полем В 1888 году Ричард Лидеккер рассмотрел «С.» Toliapicus синонимично с "C." Champsoides и "C." Ардуини , названный де Зиньо, и повторно назвал имя «С.» Спенсери для всех этих видов.

The genus name Kentisuchus was constructed only after it was realized that these specimens were clearly distinct from the genus Crocodylus and that some specimens originally assigned to "C." spenceri belonged to entirely different genera and species. "C." arduini was reassigned to the new genus Megadontosuchus in the same paper that Kentisuchus was first described in. A 2007 review of European Eocene tomistomines synonymized K. toliapicus and K. champsoides with K. spenceri.

Phylogenetics

K. spenceri skull in multiple views

K. spenceri is closely related to Megadontosuchus and Dollosuchoides. An apparent close relationship between K. spenceri and Eosuchus lerichei has been used to imply that the latter species was a tomistomine, while it is now thought that Eosuchus is a basal eusuchian that lies outside the crocodilian crown group.

Below is a cladogram based morphological studies comparing skeletal features that shows Kentisuchus as a member of Tomistominae, related to the false gharial:

Crocodylidae	Crocodylinae Tomistominae Xaymacachampsa† Megadontosuchus† Kentisuchus† Maroccosuchus† Dollosuchoides† Thecachampsa† Penghusuchus† Toyotamaphimeia† Tomistoma cairense† Maomingosuchus† Tomistoma schlegelii False gharial Gavialosuchus† Tomistoma lusitanicum† Paratomistoma† Tomistoma coppensi†

Based on morphological studies of extinct taxa, the tomistomines (including the living false gharial) were long thought to be classified as crocodiles and not closely related to gavialoids. However, recent molecular studies using DNA sequencing have consistently indicated that the false gharial (Tomistoma) (and by inference other related extinct forms in Tomistominae) actually belong to Gavialoidea (and Gavialidae).

Below is a cladogram from a 2018 tip dating study by Lee & Yates simultaneously using morphological, molecular (DNA sequencing), and stratigraphic (fossil age) data that shows Kentisuchus as a gavialoid, more basal than the last common ancestor to both the gharial and the false gharial:

Gavialoidea	Gavialidae Gavialis gangeticus Gharial Gavialis bengawanicus† Gavialis browni† Gryposuchus colombianus† Ikanogavialis† Gryposuchus pachakamue† Piscogavialis† Harpacochampsa† Toyotamaphimeia† Penghusuchus† Gavialosuchus† Tomistoma lusitanicum† Tomistoma schlegelii False gharial (crown group) Tomistoma cairense† Dollosuchoides† Maroccosuchus† Paratomistoma† Kentisuchus†
(total group)

Paleobiology

The close relation of Kentisuchus and Dollosuchoides, which are known from European localities that were on the mainland during the early Eocene, to Megadontosuchus, which is known from Italian localities that were once part of a Tethysian archipelago, suggests that it came to these islands after a dispersal event south from mainland Europe rather than north from Africa.

Внешние ссылки

• Kentisuchus в базе данных палеобиологии