Тиреофора

"Eurypoda" перенаправляет здесь. Для рода жука см. Prioninae .

Для рода мухи см. Thyreophora cynophila

Тиреофоранс Временный диапазон: ранняя юра - поздний меховой ,
Скелетная гора Гастония Буржей , Музей палеонтологии BYU
Skeletal mount of Stegosaurus stenops, Natural History Museum, London
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Clade:	Dinosauria
Clade:	†Ornithischia
Clade:	†Genasauria
Clade:	†Thyreophora Nopcsa, 1915
Subgroups
†Bienosaurus †Emausaurus †Laquintasaura? †Lesothosaurus? †Scutellosaurus †Tatisaurus †Thyreophoroidea Nopcsa, 1928 †Jakapil? †Scelidosaurus †Yuxisaurus? †Eurypoda Sereno, 1986 †Ankylosauria †Stegosauria
Thyreophorans distribution map

Thyreophora («носители щита», часто известные просто «бронированные динозавры») - это группа бронированных орнитишских динозавров , которые жили от ранней юры до конца мелового .

Тиреофораны характеризуются присутствием бронеье, выстраиваемых в продольные ряды вдоль тела. Примитивные формы имели простые, низкие, килевые бурки или остеодермы , тогда как более производные формы разработали более сложные структуры, включая шипы и пластины. Большинство тиреофоранов были травоядными и имели относительно маленький мозг для размера своего тела.

Thyreophora includes two major subgroups, Ankylosauria and Stegosauria. In both clades, the forelimbs were much shorter than the hindlimbs, particularly in stegosaurs. Thyreophora has been defined as the group consisting of all species more closely related to Ankylosaurus and Stegosaurus than to Iguanodon and Triceratops. It is the sister group of Cerapoda within Genasauria.

Characteristics

Members of Thyreophora are characterised by the presence of osteoderms (bony growths within the skin), with these osteoderms having lateral keels. Characters of the skull and jaws distinctive (synapomorphic) of thyreophorans include "absence of a deep elliptic fossa along the sutural line of the nasals, presence of a wide jugal, remodeling of skull dermal bone, down-turned dentary tooth row". Among primitive thyreophorans, Scutellosaurus was likely primarily bipedal, while the more quadrupedally adapted Scelidosaurus may have been bipedal for some of the time, particularly as a juvenile. Stegosaurs and ankylosaurs are thought to have been obligately quadrupedal.

Classification

Taxonomy

While ranked taxonomy has largely fallen out of favor among dinosaur paleontologists, a few 21st century publications have retained the use of ranks, though sources have differed on what its rank should be. Most have listed Thyreophora as an unranked taxon containing the traditional suborders Stegosauria and Ankylosauria, though Thyreophora is also sometimes classified as a suborder, with Ankylosauria and Stegosauria as infraorders.

Phylogeny

Thyreophora was first named by Nopcsa in 1915. Thyreophora was defined as a clade by Paul Sereno in 1998, as "all genasaurs more closely related to Ankylosaurus than to Triceratops". Thyreophoroidea was first named by Nopcsa in 1928 and defined by Sereno in 1986, as "Scelidosaurus, Ankylosaurus, their most recent common ancestor and all of its descendants". Eurypoda was first named by Sereno in 1986 and defined by him in 1998, as "Stegosaurus, Ankylosaurus, their most recent common ancestor and all of their descendants".

In 2021, an international group of researchers led by Daniel Madzia registered almost all of the most commonly used ornithischian clades under the International Code of Phylogenetic Nomenclature, with the intent of standardizing their definitions. According to Madzia et al., Thyreophora defined as the largest clade containing Ankylosaurus magniventris and Stegosaurus stenops but not Iguanodon bernissartensis and Triceratops horridus.

The following cladogram shows the results of the phylogenetic analysis Soto-Acuña et al. (2021). In their description of Jakapil the following year, Riguetti et al modified the same matrix and found it to occupy a position as the sister taxon to the Eurypoda. A similar result was found by Fonseca et al. in 2024.

Thyreophora	Laquintasaura Lesothosaurus Emausaurus Scutellosaurus Thyreophoroidea Scelidosauridae Scelidosaurus Yuxisaurus Jakapil Eurypoda Stegosauria Ankylosauria Parankylosauria Euankylosauria

In 2020, as part of his monograph on Scelidosaurus, David Norman revised the relationships of early thyreophorans, finding that Stegosauria was the most basal branch, with Scutellosaurus, Emausaurus and Scelidosaurus being progressive stem groups to Ankylosauria, rather than to Stegosauria+Ankylosauria. A cladogram is given below:

Thyreophora	Stegosauria   Scutellosaurus Emausaurus      Scelidosaurus Ankylosauria

See also

• thyreophoroi, Greek soldiers bearing a thyreos shield
• List of thyreophoran type specimens