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Taxonomy of Protista

A modern non-hierarchical interpretation of the phylogeny of eukaryotes, including protists

A protist (/ˈproʊtɪst/) is any eukaryotic organism (one with cells containing a nucleus) that is not an animal, plant, or fungus. The protists do not form a natural group, or clade, since they exclude certain eukaryotes with whom they share a common ancestor;[a] but, like algae or invertebrates, the grouping is used for convenience. In some systems of biological classification, such as the popular five-kingdom scheme proposed by Robert Whittaker in 1969, the protists make up a kingdom called Protista, composed of "organisms which are unicellular or unicellular-colonial and which form no tissues".[1][b] In the 21st century, the classification shifted toward a two-kingdom system of protists: Chromista (containing the chromalveolate, rhizarian and hacrobian groups) and Protozoa (containing excavates and all protists more closely related to animals and fungi).[2]

The following groups contain protists. The clade Opisthokonta also contains the animals and the fungi, and the kingdom Archaeplastida also contains algae and plants.

Key to symbols used:

(P) = Paraphyletic group.
† = Extinct group.
? = Group of uncertain taxonomic position.
* = Group lacking molecular data.
[ ] = Synonyms and alternative names.

Current classification

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This article largely follows the efforts of the scientific community and the International Society of Protistologists to revise the taxonomy of protists in a manner that reflects their phylogeny and evolution, striving away from the use of historical paraphyletic taxa and relying exclusively on clades as the basis of the classification. The reference used for the general arrangement of groups is the publication Revisions to the Classification, Nomenclature and Diversity of Eukaryotes, published in 2019 by researcher Sina Adl and coauthors.[3] Other studies are used for some specific groups of protists that have been revised after 2018, such as slime moulds,[4] glaucophyte algae,[5] arcellinid testate amoebae[6] and planktonic foraminifera.[7]

The following cladogram shows the relationships between the major groups of protists, as well as the kingdoms of animals, plants and fungi, according to the most recent phylogenetic analyses.[8][9][10][11][12]

Eukaryota
Choanozoa (P)
sensu Cavalier-Smith

Amorphea Adl et al. 2012

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Obazoa Brown et al. 2013

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Cladogram of Obazoa based on several analyses.[11][13] It includes the earliest diverging 'fungi' (microsporidians, aphelids and rozellids), which are only sometimes considered protists under the name of Opisthosporidia.[14]

Incertae sedis Obazoa:

Opisthokonta Cavalier-Smith 1987, emend. Adl et al. 2005

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Amoebozoa Lühe 1913, sensu Cavalier-Smith 1998

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Cladogram of Amoebozoa based on an 824-gene phylogenetic analysis published in 2022.[21]

Incertae sedis Amoebozoa: Belonocystis, Boveella, Biomyxa, Corallomyxa, Gibbodiscus, Hartmannia, Malamoeba, Malpighamoeba, Microglomus, Oscillosignum, Pseudothecamoeba, Rhabdamoeba, Schoutedamoeba, Stereomyxa, Subulamoeba, Thecochaos, Triaenamoeba, Unda.[3]

Tubulinea Smirnov et al. 2005

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Evosea Kang et al. 2017

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Discosea Cavalier-Smith et al. 2004 sensu Smirnov et al. 2011

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Diaphoretickes Adl et al. 2012

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Archaeplastida Adl et al. 2005

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Stramenopiles Patterson 1989, emend. Adl et al. 2005

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Stramenopiles incertae sedis:

Bigyra Cavalier-Smith 1998, emend. 2006

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Gyrista Cavalier-Smith 1998

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Alveolata Cavalier-Smith 1991

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Cladogram of Alveolata, based on a SSU rDNA phylogeny of basal alveolates[53] and a 203-gene phylogeny of apicomplexans and their relatives[54] published in 2023.

Myzozoa Cavalier-Smith & Chao 2004

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Ciliophora Doflein 1901

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Rhizaria Cavalier-Smith 2002

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Rhizaria incertae sedis: Gymnosphaerida Poche 1913, emend. Mikrjukov 2000. Actinocoryne, Cienkowskya, Gymnosphaera, Hedraiophrys [possible synonym of Cienkowskya], Wagnerella.

Cercozoa Cavalier-Smith 1998, emend. Adl et al. 2005; emend. Cavalier-Smith 2018

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Endomyxa Cavalier-Smith 2002, emend. Bass & Berney in Adl et al. 2019

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Retaria Cavalier-Smith 2002

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Haptista Cavalier-Smith 2003

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Cryptista Cavalier-Smith 1989, 2018, emend. Adl et al. 2019

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Incertae sedis Eukarya

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Hemimastigophora Foissner et al. 1988

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Excavates (P)

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Malawimonadida Cavalier-Smith 2003

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Metamonada Grassé 1952, emend. Cavalier-Smith 1987

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Discoba Simpson in Hampl et al. 2009

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CRuMs Brown et al. 2018

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Ancyromonadida Cavalier-Smith 1998

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Genera of uncertain affiliation

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Acinetactis, Actinastrum, Actinocoma, Actinolophus, Adinomonas, Aletium, Amphimonas, Amylophagus, Aphelidiopsis, Asterocaelum, Asthmatos, Aurospora, Barbetia, Berkeleyaesol,[119] Belaria, Belonocystis, Bertarellia, Bertramia, Bodopsis, Boekelovia, Branchipocola, Camptoptyche, Chalarodora, Cibdelia, Cichkovia, Cinetidomyxa, Cingula, Cladomonas, Clathrella, Codonoeca, Coelosporidium[u], Copromonas, Cyanomastix, Cyclomonas, Cytamoeba, Dallingeria, Dictyomyxa, Dimastigamoeba, Dinemula, Dinoasteromonas, Diplocalium, Diplomita, Diplophysalis, Diploselmis, Dobellina, Ducelleria, Ectobiella, Elaeorhanis, Embryocola, Endamoeba, Endemosarca, Endobiella, Endomonas, Endospora, Enteromyxa, Eperythrocytozoon, Errera, Fromentella, Gweamonas,[120] Gymnococcus, Gymnophrydium, Haematotractidium, Hartmannina, Heliobodo, Heliomonas, Hermisenella, Heterogromia, Hillea, Hyalodaktylethra, Immanoplasma, Isoselmis, Janickina, Kamera, Lagenidiopsis, Liegeosia, Luffisphaera[v], Lymphocytozoon, Lymphosporidium, Macappella, Magosphaera, Malpighiella, Martineziella, Megamoebomyxa, Meringosphaera, Microcometes, Monochrysis, Monodus, Mononema, Myrmicisporidium, Naupliicola, Nephrodinium, Neurosporidium, Orbulinella, Ovicola, Palisporomonas, Pansporella, Paradinemula, Parakaryon[w], Paraluffisphaera, Paramonas, Paraplasma, Parastasia, Parastasiella, Peliainia, Peltomonas, Petasaria, Phagodinium, Phanerobia, Phloxamoeba, Phyllomitus, Phyllomonas, Physcosporidium, Pleuophrys, Pleuromastix, Protenterospora, Protomonas, Pseudoactiniscus, Pseudosporopsis, Rhizomonas, Rhynchodinium, Rigidomastix, Schewiakoffia, Sergentella, Serpentoplasma, Sphaerasuctans, Spongastericus, Spongocyclia, Stephanomonas, Strobilomonas, Tetradimorpha, Tetragonidium, Thaulirens, Topsentella, Toshiba, Trichonema, Urbanella.[3]

See also

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Notes

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  1. ^ The first eukaryotes were “neither plants, animals, nor fungi”, hence as defined, the Protista would include the earliest common ancestor of all eukaryotes.
  2. ^ In the original 4-kingdom model proposed in 1959, Protista included all unicellular microorganisms such as bacteria.
  3. ^ Jump up to: a b This taxon belongs to the paraphyletic phylum Apusozoa Cavalier-Smith 1997 emend. 2013.[15]
  4. ^ Jump up to: a b c There are two competing phylogenetic hypotheses: Teretosporea (Corallochytrium + Ichthyosporea) and Pluriformea (Corallochytrium + Syssomonas).[3] This article follows the latter hypothesis, because it is strongly supported by the most recent phylogenetic studies.[11]
  5. ^ The term "Choanozoa" has been used since 1991 by Cavalier-Smith as a paraphyletic assemblage of opisthokont protists, and the terms "Apoikozoa" and "choanimal" were proposed as names for the clade Metazoa+Choanoflagellata. However, these terms have not been formally described or adopted, and were rejected in favour of a renamed Choanozoa to fit the clade Metazoa+Choanoflagellata.[3]
  6. ^ The position of the genera Microcorycia, Parmulina, Penardochlamys and Zonomyxa, which were listed in 2002 under family Microcoryciidae, is not clear. They are placed here by morphological characters but this needs to be supported by molecular data.[3]
  7. ^ The latest revision by Adl et al. (2019) ignores the grouping of some variosean genera into higher rank clades that was made in older studies because of the weakly supported SSU phylogenetic analyses, except for Fractovitellida, which is fully supported by phylogenomic analyses from 2017.[3]
  8. ^ Jump up to: a b c d e f g This order was initially divided into families, but phylogenetic analyses haven't supported the monophyly of each family. For this reason, in recent classifications there is no family rank division within it.[22][3]
  9. ^ This genus groups with Dermamoebidae in a phylogenetic anaysis that uses a limited number of taxa.[30] In the recent taxonomy it is listed as a separate clade until its position is better resolved.[3]
  10. ^ This genus could be a synonym of Vannella.[3]
  11. ^ Both genera group together in some phylogenetic analyses, but usually appear separated. They are considered a potential branch of Discosea, unless proven otherwise in the future with better taxon sampling.[3]
  12. ^ Latest analyses show that Microheliella is the sister group to Cryptista.[12]
  13. ^ Telonemia is the sister group to the SAR clade.[9]
  14. ^ The picobiliphytes are phylogenetically closer to Rhodophyta and Rhodelphis than to other groups, and are therefore considered part of Archaeplastida.[36]
  15. ^ Cavalier-Smith and Scoble (2013) show a sister relationship between raphidophytes and Actinophryida.[47]
  16. ^ Apart from morphological descriptions, no molecular analysis has solidly affiliated Rastrimonas with the Perkinsozoa.[58]
  17. ^ Highly divergent 18S rRNA.[3]
  18. ^ This taxon is artificial; many of the families listed in it are not monophyletic and have little support from phylogenetic analyses.[3]
  19. ^ There is only one molecular sequence from this genus, which causes long branches in gene phylogenies, and the node is unresolved.[3]
  20. ^ According to Nakamura & Suzuki, the only extant (i.e. not extinct) orders of Polycystinea are Spumellaria, Nassellaria, Collodaria and Orodaria. Therefore, Entactinaria is considered an exclusively fossil group.[95]
  21. ^ Probably a synonym of the zygomycete fungus Nephridiophaga.[3]
  22. ^ May be the same genus as Belonocystis.[3]
  23. ^ It is not yet known whether this organism is an eukaryote or a prokaryote.[121]

References

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  1. ^ Jump up to: a b Whittaker, R. H. (1969). "New Concepts of Kingdoms of Organisms". Science. 163 (3863): 150–160. Bibcode:1969Sci...163..150W. doi:10.1126/SCIENCE.163.3863.150. PMID 5762760.
  2. ^ Ruggiero, Michael A.; Gordon, Dennis P.; Orrell, Thomas M.; Bailly, Nicolas; Bourgoin, Thierry; Brusca, Richard C.; Cavalier-Smith, Thomas; Guiry, Michael D.; Kirk, Paul M. (2015). "A Higher Level Classification of All Living Organisms". PLOS ONE. 10 (4): e0119248. Bibcode:2015PLoSO..1019248R. doi:10.1371/JOURNAL.PONE.0119248. PMC 4418965. PMID 25923521. (Erratum: doi:10.1371/journal.pone.0130114)
  3. ^ Jump up to: a b c d e f g h i j k l m n o p q Adl, Sina M.; et al. (2019). "Revisions to the Classification, Nomenclature, and Diversity of Eukaryotes". Journal of Eukaryotic Microbiology. 66 (1): 4–119. doi:10.1111/JEU.12691. PMC 6492006. PMID 30257078.
  4. ^ Jump up to: a b c d e f g h i j k Leontyev, Dmitry V.; Schnittler, Martin; Stephenson, Steven L.; Novozhilov, Yuri K.; Shchepin, Oleg N. (2019). "Towards a phylogenetic classification of the Myxomycetes". Phytotaxa. 399 (3): 209. doi:10.11646/PHYTOTAXA.399.3.5.
  5. ^ Jump up to: a b Figueroa-Martinez F, Jackson C, Reyes-Prieto R (January 2019). "Plastid Genomes from Diverse Glaucophyte Genera Reveal a Largely Conserved Gene Content and Limited Architectural Diversity". Genome Biology and Evolution. 11 (1): 174–188. doi:10.1093/gbe/evy268. PMC 6330054. PMID 30534986.
  6. ^ Jump up to: a b c d González-Miguéns, Rubén; Todorov, Milcho; Blandenier, Quentin; Duckert, Clément; Porfirio-Sousa, Alfredo L.; Ribeiro, Giulia M.; Ramos, Diana; Lahr, Daniel J.G.; Buckley, David; Lara, Enrique (2022). "Deconstructing Difflugia: The tangled evolution of lobose testate amoebae shells (Amoebozoa: Arcellinida) illustrates the importance of convergent evolution in protist phylogeny". Molecular Phylogenetics and Evolution. 175. Bibcode:2022MolPE.17507557G. doi:10.1016/J.YMPEV.2022.107557. hdl:10261/281619. PMID 35777650.
  7. ^ Jump up to: a b c d e f g Arenillas I, Arz JA, Gilabert V (2022). "An updated suprageneric classification of planktic foraminifera after growing evidence of multiple benthic-planktic transitions". Spanish Journal of Palaeontology. 37 (1). Sociedad Española de Paleontología: 1–34. doi:10.7203/sjp.22189. ISSN 2255-0550.
  8. ^ Derelle R, Torruella G, Klimeš V, Brinkmann H, Kim E, Vlček Č, Lang BF, Eliáš M (2015), "Bacterial proteins pinpoint a single eukaryotic root", Proceedings of the National Academy of Sciences, 112 (7): E693–699, Bibcode:2015PNAS..112E.693D, doi:10.1073/pnas.1420657112, PMC 4343179, PMID 25646484
  9. ^ Jump up to: a b Brown MW, et al. (2018), "Phylogenomics Places Orphan Protistan Lineages in a Novel Eukaryotic Super-Group", Genome Biology and Evolution, 10 (2): 427–433, doi:10.1093/gbe/evy014, PMC 5793813, PMID 29360967
  10. ^ Jump up to: a b Tikhonenkov, Denis V.; Mikhailov, Kirill V.; Gawryluk, Ryan M. R.; Belyaev, Artem O.; Mathur, Varsha; Karpov, Sergey A.; Zagumyonnyi, Dmitry G.; Borodina, Anastasia S.; Prokina, Kristina I.; Mylnikov, Alexander P.; Aleoshin, Vladimir V.; Keeling, Patrick J. (2022). "Microbial predators form a new supergroup of eukaryotes". Nature. 612 (7941): 714–719. Bibcode:2022Natur.612..714T. doi:10.1038/s41586-022-05511-5. PMID 36477531. S2CID 254436650.
  11. ^ Jump up to: a b c d Tikhonenkov, Denis V.; Mikhailov, Kirill V.; Hehenberger, Elisabeth; Karpov, Sergei A.; Prokina, Kristina I.; Esaulov, Anton S.; Belyakova, Olga I.; Mazei, Yuri A.; Mylnikov, Alexander P.; Aleoshin, Vladimir V.; Keeling, Patrick J. (2020). "New Lineage of Microbial Predators Adds Complexity to Reconstructing the Evolutionary Origin of Animals". Current Biology. 30 (22): 4500–4509.e5. Bibcode:2020CBio...30E4500T. doi:10.1016/J.CUB.2020.08.061. PMID 32976804.
  12. ^ Jump up to: a b Euki, Yazaki; Akinori, Yabuki; Ayaka, Imaizumi; Keitaro, Kume; Tetsuo, Hashimoto; Yuji, Inagaki (2022). "The closest lineage of Archaeplastida is revealed by phylogenomics analyses that include Microheliella maris". Open Biology. 12 (4): 210376. doi:10.1098/rsob.210376. PMC 9006020. PMID 35414259.
  13. ^ Galindo LJ, Torruella G, López-García P, Ciobanu M, Gutiérrez-Preciado A, Karpov SA, Moreira D (2023). "Phylogenomics supports the monophyly of aphelids and fungi and identifies new molecular synapomorphies". Systematic Biology. 72 (3): 505–515. doi:10.1093/sysbio/syac054. PMID 35900180.
  14. ^ Wijayawardene, NN; Hyde, KD; Al-Ani, LKT; Tedersoo, L; Haelewaters, D; Rajeshkumar, KC; Zhao, RL (18 March 2020). "Outline of Fungi and fungus-like taxa". Mycosphere. 11 (1: 8): 1060–1456. doi:10.5943/mycosphere/11/1/8. hdl:10481/61998. ISSN 2077-7019.
  15. ^ Cavalier-Smith T (December 2021). "Ciliary transition zone evolution and the root of the eukaryote tree: implications for opisthokont origin and classification of kingdoms Protozoa, Plantae, and Fungi". Protoplasma. 259 (3): 487–593. doi:10.1007/s00709-021-01665-7. PMC 9010356. PMID 34940909.
  16. ^ Torruella G, Galindo LJ, Moreira D, Ciobanu M, Heiss AA, Yubuki N, et al. (2022). "Expanding the molecular and morphological diversity of Apusomonadida, a deep-branching group of gliding bacterivorous protists". Journal of Eukaryotic Microbiology. 70 (2): e12956. doi:10.1111/jeu.12956. hdl:2117/404026. PMID 36453005. S2CID 253460648.
  17. ^ Gabaldón T, Völcker E, Torruella G (2022). "On the Biology, Diversity and Evolution of Nucleariid Amoebae (Amorphea, Obazoa, Opisthokonta". Protist. 173 (4): 125895. doi:10.1016/j.protis.2022.125895. hdl:2117/369912. PMID 35841659. S2CID 249827842.
  18. ^ Strother, Paul K.; Brasier, Martin D.; Wacey, David; Timpe, Leslie; Saunders, Martin; Wellman, Charles H. (2021). "A possible billion-year-old holozoan with differentiated multicellularity". Current Biology. 31 (12): 2658–2665.e2. Bibcode:2021CBio...31E2658S. doi:10.1016/J.CUB.2021.03.051. PMID 33852871.
  19. ^ Urrutia A, Mitsi K, Foster R, Ross S, Carr M, Ward GM, et al. (2022). "Txikispora philomaios n. sp., n. g., a micro-eukaryotic pathogen of amphipods, reveals parasitism and hidden diversity in Class Filasterea". Journal of Eukaryotic Microbiology. 69 (2): e12875. doi:10.1111/jeu.12875. PMID 34726818. S2CID 240422937.
  20. ^ Schiwitza, Sabine; Arndt, Hartmut; Nitsche, Frank (2019). "First description of an euryoecious acanthoecid choanoflagellate species, Enibas tolerabilis gen. et sp. nov. from a salar in the Chilean Andes based on morphological and transcriptomic data". European Journal of Protistology. 67: 106–113. doi:10.1016/j.ejop.2018.11.004. PMID 30572146. S2CID 58552347.
  21. ^ Tekle YI, Wang F, Wood FC, Anderson OR, Smirnov A (2022). "New insights on the evolutionary relationships between the major lineages of Amoebozoa". Scientific Reports. 12 (11173): 11173. Bibcode:2022NatSR..1211173T. doi:10.1038/s41598-022-15372-7. PMC 9249873. PMID 35778543. S2CID 247231712.
  22. ^ Jump up to: a b c Kang, Seungho; Tice, Alexander K; Spiegel, Frederick W; Silberman, Jeffrey D; Pánek, Tomáš; Čepička, Ivan; Kostka, Martin; Kosakyan, Anush; Alcântara, Daniel M C; Roger, Andrew J; Shadwick, Lora L; Smirnov, Alexey; Kudryavtsev, Alexander; Lahr, Daniel J G; Brown, Matthew W (September 2017). "Between a Pod and a Hard Test: The Deep Evolution of Amoebae". Molecular Biology and Evolution. 34 (9): 2258–2270. doi:10.1093/molbev/msx162. PMC 5850466. PMID 28505375.
  23. ^ Jump up to: a b Smirnov, Alexey; Nassonova, Elena; Geisen, Stefan; Bonkowski, Michael; Kudryavtsev, Alexander; Berney, Cedric; Glotova, Anna; Bondarenko, Natalya; Dyková, Iva; Mrva, Martin; Fahrni, Jose; Pawlowski, Jan (2017). "Phylogeny and Systematics of Leptomyxid Amoebae (Amoebozoa, Tubulinea, Leptomyxida)". Protist. 168 (2): 220–252. doi:10.1016/j.protis.2016.10.006. ISSN 1434-4610. PMID 28343121.
  24. ^ Vašíček, Miloslav; Růžička, Bohuslav (1957). "Namurian Thecamoebina from the Ostrava-Karviná Coal District". Acta Musei Nationalis Pragae. 13B (5): 333–340.
  25. ^ Bobrov, Anatoly; Mazei, Yuri (2017). "A review of testate amoeba genus Cryptodifflugia Penard, 1890 (Phryganellina: Cryptodifflugiidae) with a key to species". Zootaxa. 4282 (2): 292–308. doi:10.11646/zootaxa.4282.2.4.
  26. ^ Jump up to: a b c Cavalier-Smith, Thomas; Chao, Ema E.; Lewis, Rhodri (2016). "187-gene phylogeny of protozoan phylum Amoebozoa reveals a new class (Cutosea) of deep-branching, ultrastructurally unique, enveloped marine Lobosa and clarifies amoeba evolution". Molecular Phylogenetics and Evolution. 99: 275–296. Bibcode:2016MolPE..99..275C. doi:10.1016/J.YMPEV.2016.03.023. PMID 27001604.
  27. ^ Lahr, Daniel J. G.; Grant, Jessica; Nguyen, Truc; Lin, Jian Hua; Katz, Laura A. (28 July 2011). "Comprehensive Phylogenetic Reconstruction of Amoebozoa Based on Concatenated Analyses of SSU-rDNA and Actin Genes". PLOS ONE. 6 (7): e22780. Bibcode:2011PLoSO...622780L. doi:10.1371/JOURNAL.PONE.0022780. PMC 3145751. PMID 21829512.
  28. ^ Lloyd SJ, Leontyev DV, Moreno G, Villalba AL, Schnittler M (2023). "Tasmaniomyxa umbilicata, a new genus and new species of myxomycete from Tasmania". Mycologia. 116 (1): 170–183. doi:10.1080/00275514.2023.2274252. PMID 38032605. S2CID 265504848.
  29. ^ Hess S, Eme L, Roger AJ, Simpson AG (2019). "A natural toroidal microswimmer with a rotary eukaryotic flagellum". Nature Microbiology. 4 (10): 1620–1626. doi:10.1038/s41564-019-0478-6. PMID 31182800. S2CID 182951779.
  30. ^ Blandenier Q, Seppey CVW, Singer D, Vlimant M, Simon A, Duckert C, Lara E (2016). "Mycamoeba gemmipara nov. gen., nov. sp., the First Cultured Member of the Environmental Dermamoebidae Clade LKM74 and its Unusual Life Cycle" (PDF). Journal of Eukaryotic Microbiology. 64 (2): 257–265. doi:10.1111/jeu.12357. PMID 27543384. S2CID 4665573.
  31. ^ Kudryavtsev, Alexander; Voytinsky, Fyodor; Volkova, Ekaterina (July 2022). "Coronamoeba villafranca gen. nov. sp. nov. (Amoebozoa, Dermamoebida) challenges the correlation of morphology and phylogeny in Amoebozoa". Scientific Reports. 12 (1): 12541. Bibcode:2022NatSR..1212541K. doi:10.1038/s41598-022-16721-2. PMC 9307759. PMID 35869259.
  32. ^ Melton JT, Wood FC, Branch J, Singla M, Tekle YI (2019). "Phylogenomics of Thecamoebida (Discosea, Amoebozoa) with the Description of Stratorugosa tubuloviscum gen. nov. sp. nov., a Freshwater Amoeba with a Perinuclear MTOC". Protist. 170 (1): 8–20. doi:10.1016/j.protis.2018.09.002. ISSN 1434-4610. PMID 30553127. S2CID 56174714.
  33. ^ Mesentsev Y, Bondarenko N, Kamyshatskaya O, Nassonova E, Glotova A, Loiko S, Istigechev G, Kulemzina A, Abakumov E, Rayko M, Lapidus A, Smirnov A (2023). "Thecochaos is not a myth: study of the genus Thecochaos (Amoebozoa, Discosea) – a rediscovered group of lobose amoeba, with short SSU gene". Organisms Diversity & Evolution. 23: 7–24. doi:10.1007/s13127-022-00581-9. S2CID 251658030.
  34. ^ Tice AK, Shadwick LL, Fiore-Donno AM, et al. (2016). "Expansion of the molecular and morphological diversity of Acanthamoebidae (Centramoebida, Amoebozoa) and identification of a novel life cycle type within the group". Biology Direct. 11 (1): 69. doi:10.1186/s13062-016-0171-0. PMC 5192571. PMID 28031045.
  35. ^ Tikhonenkov DV, Jamy M, Borodina AS, Belyaev AO, Zagumyonnyi DG, Prokina KI, Mylnikov AP, Burki F, Karpov SA (2022). "On the origin of TSAR: morphology, diversity and phylogeny of Telonemia". Open Biology. 12 (3): 210325. doi:10.1098/rsob.210325. PMC 8924772. PMID 35291881.
  36. ^ Schön ME, Zlatogursky VV, Singh RP, et al. (2021). "Single cell genomics reveals plastid-lacking Picozoa are close relatives of red algae". Nature Communications. 12 (1): 6651. Bibcode:2021NatCo..12.6651S. doi:10.1038/s41467-021-26918-0. PMC 8599508. PMID 34789758.
  37. ^ Gawryluk, Ryan M. R.; Tikhonenkov, Denis V.; Hehenberger, Elisabeth; Husnik, Filip; Mylnikov, Alexander P.; Keeling, Patrick J. (August 2019). "Non-photosynthetic predators are sister to red algae". Nature. 572 (7768): 240–243. doi:10.1038/s41586-019-1398-6. ISSN 1476-4687. PMID 31316212. S2CID 197542583.
  38. ^ Weston EJ, Eglit Y, Simpson AG (2023). "Kaonashia insperata gen. et sp. nov., a eukaryotrophic flagellate, represents a novel major lineage of heterotrophic stramenopiles". Journal of Eukaryotic Microbiology. 71 (1): e13003. doi:10.1111/jeu.13003. PMID 37803921.
  39. ^ Rybarski AE, Nitsche F, Park JS, Filz P, Schmidt P, Kondo R, Simpson AG, Arndt H (2021). "Revision of the phylogeny of Placididea (Stramenopiles): Molecular and morphological diversity of novel placidid protists from extreme aquatic environments". European Journal of Protistology. 81: 125809. doi:10.1016/j.ejop.2021.125809. ISSN 0932-4739. PMID 34673437. S2CID 236377023.
  40. ^ Jirsová D, Füssy Z, Richtová J, Gruber A, Oborník M (2019). "Morphology, Ultrastructure, and Mitochondrial Genome of the Marine Non-Photosynthetic Bicosoecid Cafileria marina Gen. et sp. nov". Microorganisms. 7 (8): 240. doi:10.3390/microorganisms7080240. PMC 6723347. PMID 31387253.
  41. ^ Harder, Christoffer Bugge; Ekelund, Flemming; Karpov, Sergey A. (March 2014). "Ultrastructure and phylogenetic position of Regin rotiferus and Otto terricolus genera et species novae (Bicosoecida, Heterokonta/Stramenopiles)". Protist. 165 (2): 144–160. doi:10.1016/j.protis.2014.01.004. PMID 24637333.
  42. ^ Hassett, Brandon T. (2020). "A Widely Distributed Thraustochytrid Parasite of Diatoms Isolated from the Arctic Represents a gen. and sp. nov". Journal of Eukaryotic Microbiology. 67 (4). International Society of Protistologists: 480–490. doi:10.1111/jeu.12796. hdl:10037/19021. PMID 32249965.
  43. ^ Shiratori, Takashi; Thakur, Rabindra; Ishida, Ken-ichiro (2017). "Pseudophyllomitus vesiculosus (Larsen and Patterson 1990) Lee, 2002, a Poorly Studied Phagotrophic Biflagellate is the First Characterized Member of Stramenopile Environmental Clade MAST-6". Protist. 168 (4): 439–451. doi:10.1016/j.protis.2017.06.004. ISSN 1434-4610. PMID 28822908.
  44. ^ Dovilė Barcytė; Wenche Eikrem; Anette Engesmo; Sergio Seoane; Jens Wohlmann; Aleš Horák; Tatiana Yurchenko; Marek Eliáš (2 March 2021). "Olisthodiscus represents a new class of Ochrophyta". Journal of Phycology. 57 (4): 1094–1118. Bibcode:2021JPcgy..57.1094B. doi:10.1111/jpy.13155. hdl:10852/86515. PMID 33655496.
  45. ^ Amaral R, Fawley KP, Němcová Y, Ševčíková T, Lukešová A, Fawley MW, Santos LM, Eliáš M (2020). "Toward Modern Classification of Eustigmatophytes, Including the Description of Neomonodaceae Fam. Nov. and Three New Genera". Journal of Phycology. 56 (3): 630–648. Bibcode:2020JPcgy..56..630A. doi:10.1111/jpy.12980. PMC 7987219. PMID 32068883.
  46. ^ Cavalier-Smith, Thomas (2017). "Kingdom Chromista and its eight phyla: a new synthesis emphasising periplastid protein targeting, cytoskeletal and periplastid evolution, and ancient divergences". Protoplasma. 255 (1): 297–357. doi:10.1007/s00709-017-1147-3. PMC 5756292. PMID 28875267.
  47. ^ Cavalier-Smith, Thomas; Scoble, Josephine Margaret (2013). "Phylogeny of Heterokonta: Incisomonas marina, a uniciliate gliding opalozoan related to Solenicola (Nanomonadea), and evidence that Actinophryida evolved from raphidophytes". European Journal of Protistology. 49 (3): 328–353. doi:10.1016/j.ejop.2012.09.002. PMID 23219323.
  48. ^ Wetherbee R, Jackson CJ, Repetti SI, Clementson LA, Costa JF, van de Meene A, Crawford S, Verbruggen H (April 2019). "The golden paradox - a new heterokont lineage with chloroplasts surrounded by two membranes". Journal of Phycology. 55 (2): 257–278. Bibcode:2019JPcgy..55..257W. doi:10.1111/jpy.12822. hdl:11343/233613. PMID 30536815. S2CID 54477112.
  49. ^ Graf L, Yang EC, Han KY, Küpper FC, Benes KM, Oyadomari JK, Herbert RJH, Verbruggen H, Wetherbee R, Andersen RA, Yoon HS (December 2020). "Multigene Phylogeny, Morphological Observation and Re-examination of the Literature Lead to the Description of the Phaeosacciophyceae Classis Nova and Four New Species of the Heterokontophyta SI Clade". Protist. 171 (6): 125781. doi:10.1016/j.protis.2020.125781. PMID 33278705. S2CID 227315556.
  50. ^ Kulikovskiy M, Maltsev Y, Andreeva S, Glushchenko A, Gusev E, Podunay Y, Ludwig TV, Tusset E, Kociolek JP (2019). "Description of a new diatom genus Dorofeyukea gen. nov. with remarks on phylogeny of the family Stauroneidaceae". Journal of Phycology. 55 (1): 173–185. Bibcode:2019JPcgy..55..173K. doi:10.1111/jpy.12810. PMID 30379324.
  51. ^ Bente Edvardsen; Wenche Eikrem; Kamran Shalchian-Tabrizi; Ingvild Riisberg; Geir Johnsen; Lars Naustvoll; Jahn Throndsen (15 September 2007). "Verrucophora farcimen gen. et sp. nov. (Dictyochophyceae, Heterokonta)—a bloom-forming ichthyotoxic flagellate from the Skagerrak, Norway". Journal of Phycology. 43 (5): 1054–1070. Bibcode:2007JPcgy..43.1054E. doi:10.1111/j.1529-8817.2007.00390.x. hdl:10852/11590.
  52. ^ Han, Kwi Young; Graf, Louis; Reyes, Carolina P.; Melkonian, Barbara; Andersen, Robert A.; Yoon, Hwan Su; Melkonian, Michael (2018). "A Re-investigation of Sarcinochrysis marina (Sarcinochrysidales, Pelagophyceae) from its Type Locality and the Descriptions of Arachnochrysis, Pelagospilus, Sargassococcus and Sungminbooa genera nov". Protist. 169 (1): 79–106. doi:10.1016/j.protis.2017.12.004. ISSN 1434-4610. PMID 29427838.
  53. ^ Jump up to: a b c Gigeroff AS, Eglit Y, Simpson AG (2023). "Characterisation and cultivation of new lineages of colponemids, a critical assemblage for inferring alveolate evolution". Protist. 174 (2): 125949. doi:10.1016/j.protis.2023.125949. PMID 37019068. S2CID 257625613.
  54. ^ Mathur, Varsha; Salomaki, Eric D.; Wakeman, Kevin C.; Na, Ina; Kwong, Waldan K.; Kolisko, Martin; Keeling, Patrick J. (2023). "Reconstruction of Plastid Proteomes of Apicomplexans and Close Relatives Reveals the Major Evolutionary Outcomes of Cryptic Plastids". Molecular Biology and Evolution. doi:10.1093/MOLBEV/MSAD002. PMC 9847631. PMID 36610734.
  55. ^ Janouškovec, Jan; Tikhonenkov, Denis V.; Burki, Fabien; Howe, Alexis T.; Kolísko, Martin; Mylnikov, Alexander P.; Keeling, Patrick J. (2015). "Factors mediating plastid dependency and the origins of parasitism in apicomplexans and their close relatives". Proceedings of the National Academy of Sciences. 112 (33): 10200–10207. Bibcode:2015PNAS..11210200J. doi:10.1073/PNAS.1423790112. PMC 4547307. PMID 25717057.
  56. ^ Guiry, Michael D. (2024). "How many species of algae are there? A reprise. Four kingdoms, 14 phyla, 63 classes and still growing". Journal of Phycology. 60 (2): 214–228. Bibcode:2024JPcgy..60..214G. doi:10.1111/JPY.13431. PMID 38245909.
  57. ^ Mathur, Varsha; Kolísko, Martin; Hehenberger, Elisabeth; Irwin, Nicholas A.T.; Leander, Brian S.; Kristmundsson, Árni; Freeman, Mark A.; Keeling, Patrick J. (2019). "Multiple Independent Origins of Apicomplexan-Like Parasites". Current Biology. 29 (17): 2936–2941.e5. Bibcode:2019CBio...29E2936M. doi:10.1016/J.CUB.2019.07.019. PMID 31422883.
  58. ^ Itoïz, Sarah; Metz, Sebastian; Derelle, Evelyne; Reñé, Albert; Garcés, Esther; Bass, David; Soudant, Philippe; Chambouvet, Aurélie (2022). "Emerging Parasitic Protists: The Case of Perkinsea". Frontiers in Microbiology. 12. doi:10.3389/FMICB.2021.735815. PMC 8792838. PMID 35095782.
  59. ^ Reñé, Albert; Alacid, Elisabet; Gallisai, Rachele; Chambouvet, Aurélie; Fernández-Valero, Alan D.; Garcés, Esther (2021). "New Perkinsea Parasitoids of Dinoflagellates Distantly Related to Parviluciferaceae Members". Frontiers in Microbiology. 12. doi:10.3389/FMICB.2021.701196. PMC 8375308. PMID 34421856.
  60. ^ Jeon, Boo Seong; Park, Myung Gil (2021). "A Novel Parasitoid of Marine Dinoflagellates, Pararosarium dinoexitiosum gen. Et sp. Nov. (Perkinsozoa, Alveolata), Showing Characteristic Beaded Sporocytes". Frontiers in Microbiology. 12. doi:10.3389/FMICB.2021.748092. PMC 8667275. PMID 34912310.
  61. ^ Reñé, Albert; Alacid, Elisabet; Ferrera, Isabel; Garcés, Esther (2017). "Evolutionary Trends of Perkinsozoa (Alveolata) Characters Based on Observations of Two New Genera of Parasitoids of dinoflagellates, Dinovorax gen. nov. And snorkelia gen. nov". Frontiers in Microbiology. 8: 1594. doi:10.3389/FMICB.2017.01594. PMC 5609580. PMID 28970818.
  62. ^ Jeon, Boo Seong; Park, Myung Gil (2019). "Tuberlatum coatsi gen. n., sp. n. (Alveolata, Perkinsozoa), a New Parasitoid with Short Germ Tubes Infecting Marine Dinoflagellates". Protist. 170 (1): 82–103. doi:10.1016/J.PROTIS.2018.12.003. PMID 30797136.
  63. ^ Evans, Clive W.; Patel, Selina; Matzke, Nicholas J.; Millar, Craig D. (2023). "Cryoxcellia borchgrevinki gen. nov., sp. nov., a new parasitic X-cell species in an Antarctic nototheniid fish, the bald notothen Trematomus borchgrevinki". Polar Biology. 46 (6): 513–521. Bibcode:2023PoBio..46..513E. doi:10.1007/S00300-023-03132-W.
  64. ^ Desvignes, Thomas; Lauridsen, Henrik; Valdivieso, Alejandro; Fontenele, Rafaela S.; Kraberger, Simona; Murray, Katrina N.; Le François, Nathalie R.; Detrich, H. William; Kent, Michael L.; Varsani, Arvind; Postlethwait, John H. (2022). "A parasite outbreak in notothenioid fish in an Antarctic fjord". iScience. 25 (7). Bibcode:2022iSci...25j4588D. doi:10.1016/J.ISCI.2022.104588. PMC 9253362. PMID 35800770.
  65. ^ Karlsbakk, Egil; Nystøyl, Cecilie Flatnes; Plarre, Heidrun; Nylund, Are (2021). "A novel protist parasite, Salmoxcellia vastator n. gen., n. sp. (Xcelliidae, Perkinsozoa), infecting farmed salmonids in Norway". Parasites & Vectors. 14 (1): 431. doi:10.1186/S13071-021-04886-0. PMC 8400403. PMID 34454593.
  66. ^ Okamoto N, Horák A, Keeling PJ (2012). "Description of Two Species of Early Branching Dinoflagellates, Psammosa pacifica n. g., n. sp. and P. atlantica n. sp". PLOS ONE. 7 (6): e34900. Bibcode:2012PLoSO...734900O. doi:10.1371/journal.pone.0034900. PMC 3377698. PMID 22719825.
  67. ^ Jump up to: a b c d Holt CC, Hehenberger E, Tikhonenkov DV, Jacko-Reynolds VK, Okamoto N, Cooney EC, Irwin NA, Keeling PJ (2023). "Multiple parallel origins of parasitic Marine Alveolates". Nature. 14 (1): 7049. Bibcode:2023NatCo..14.7049H. doi:10.1038/s41467-023-42807-0. PMC 10624901. PMID 37923716.
  68. ^ Yoo J, Kim S, Wayne Coats D (2024). "Morphology of Hobagella saltata n. gen. and n. sp. (Syndiniophyceae, Miozoa) infecting the marine dinoflagellate Cucumeridinium coeruleum (Dinophyceae, Miozoa) and its potential onshore advection". Frontiers in Marine Science. 10: 1296836. doi:10.3389/fmars.2023.1296836.
  69. ^ Luo Z, Hu Z, Tang Y, Mertens KN, Leaw CP, Lim PT, Teng ST, Wang L, Gu H (2018). "Morphology, ultrastructure, and molecular phylogeny of Wangodinium sinense gen. et sp. nov. (Gymnodiniales, Dinophyceae) and revisiting of Gymnodinium dorsalisulcum and Gymnodinium impudicum" (PDF). Journal of Phycology. 54 (5): 744–761. Bibcode:2018JPcgy..54..744L. doi:10.1111/jpy.12780. PMID 30144373. S2CID 52082067.
  70. ^ Saburova M, Chomérat N (2019). "Laciniporus arabicus gen. et sp. nov. (Dinophyceae, Peridiniales), a new thecate, marine, sand-dwelling dinoflagellate from the northern Indian Ocean (Arabian Sea)" (PDF). Journal of Phycology. 55 (1): 84–103. Bibcode:2019JPcgy..55...84S. doi:10.1111/jpy.12783. PMID 30179255. S2CID 206148081.
  71. ^ Mathur, Varsha; Wakeman, Kevin C.; Keeling, Patrick J. (2021). "Parallel functional reduction in the mitochondria of apicomplexan parasites". Current Biology. 31 (13): 2920–2928.e4. Bibcode:2021CBio...31E2920M. doi:10.1016/J.CUB.2021.04.028. PMID 33974849.
  72. ^ Varsha Mathur (November 2020). The phylogeny and evolution apicomplexan parasites (PhD thesis). Vancouver: University of British Columbia. doi:10.14288/1.0395117.
  73. ^ Weiss, James; Andreou, Demetra; Esteban, Genoveva F. (2022). "The Extraordinarily Rare Ciliate Legendrea loyezae Fauré-Fremiet, 1908 (Haptoria, Ciliophora)". Protist. 173 (6): 125912. doi:10.1016/j.protis.2022.125912. PMID 36242851.
  74. ^ Ito, Akira; Ishihara, Miki; Imai, Soichi (2014). "Bozasella gracilis n. sp. (Ciliophora, Entodiniomorphida) from Asian elephant and phylogenetic analysis of entodiniomorphids and vestibuliferids". European Journal of Protistology. 50 (2): 134–152. doi:10.1016/J.EJOP.2014.01.003. PMID 24703615.
  75. ^ Irwin, Nicholas A.T.; Tikhonenkov, Denis V.; Hehenberger, Elisabeth; Mylnikov, Alexander P.; Burki, Fabien; Keeling, Patrick J. (2019-01-01). "Phylogenomics supports the monophyly of the Cercozoa". Molecular Phylogenetics and Evolution. 130: 416–423. Bibcode:2019MolPE.130..416I. doi:10.1016/j.ympev.2018.09.004. ISSN 1055-7903. PMID 30318266. S2CID 52982396.
  76. ^ Feng, Jian-Ju; He, Chen-Yang; Jiang, Shu-Hua; Zhang, Tao; Yu, Li-Yan (2021). "Saccharomycomorpha psychra n. g., n. sp., a Novel Member of Glissmonadida (Cercozoa) Isolated from Arctic and Antarctica". Journal of Eukaryotic Microbiology. 68 (3): e12840. doi:10.1111/jeu.12840. PMID 33448091.
  77. ^ Dumack K, Pundt J, Bonkowski M (2020). "Transfer of the Thecate Amoeba Lecythium mutabilis to a Novel Genus Omnivora (Fiscullidae, Thecofilosea, Cercozoa)". Journal of Eukaryotic Microbiology. 67 (2): 245–251. doi:10.1111/jeu.12778. PMID 31808200. S2CID 208768245.
  78. ^ Dumack K, Duckert C, Meinhardt R, Lara E, Bonkowski M (2021). "Description of Phaeobola aeris gen. nov., sp. nov (Rhizaria, Cercozoa, Euglyphida) Sheds Light on Euglyphida's Dark Matter". Journal of Eukaryotic Microbiology. 68 (2): e12835. doi:10.1111/jeu.12835. hdl:10261/255052. PMID 33222324. S2CID 227135214.
  79. ^ Soler-Zamora, Carmen; Useros, Fernando; González-Miguéns, Rubén; Gómez-Rodríguez, Pablo; Lara, Enrique (10 July 2023). "The problem of 'shadow species' as illustrated with the taxonomic hotchpotch Cyphoderia ampulla (Rhizaria: Cyphoderiidae)". Zoological Journal of the Linnean Society. XX: 1–16. doi:10.1093/zoolinnean/zlad040.
  80. ^ Siemensma F, Dumack K (2020). "SSU rDNA Phylogeny Indicates the Scale-lacking Trivalvulariida ord. nov. as a Sister Group to the Euglyphida (Cercozoa, Rhizaria)". Protist. 171 (1): 125701. doi:10.1016/j.protis.2019.125701. ISSN 1434-4610. PMID 31955058. S2CID 210829787.
  81. ^ Bass D, Chao EE, Nikolaev S, Yabuki A, Ishida KI, Berney C, Pakzad U, Wylezich C, Cavalier-Smith T (2009). "Phylogeny of Novel Naked Filose and Reticulose Cercozoa: Granofilosea cl. n. and Proteomyxidea Revised". Protist. 160 (1): 75–109. doi:10.1016/j.protis.2008.07.002. ISSN 1434-4610. PMID 18952499.
  82. ^ Hess S, Suthaus A (2022). "The Vampyrellid Amoebae (Vampyrellida, Rhizaria)". Protist. 173 (1): 125854. doi:10.1016/j.protis.2021.125854. PMID 35091168. S2CID 245303468.
  83. ^ Suthaus A, Hess S (2023). "Pseudovampyrella gen. nov.: A genus of Vampyrella-like protoplast extractors finds its place in the Leptophryidae". The Journal of Eukaryotic Microbiology. 71 (1). International Society of Protistologists: e13002. doi:10.1111/jeu.13002. PMID 37743754.
  84. ^ Kolátková V, Čepička I, Hoffman R, et al. (2021). "Marinomyxa Gen. Nov. Accommodates Gall-Forming Parasites of the Tropical to Subtropical Seagrass Genus Halophila and Constitutes a Novel Deep-Branching Lineage Within Phytomyxea (Rhizaria: Endomyxa)". Microbial Ecology. 81 (3): 673–686. Bibcode:2021MicEc..81..673K. doi:10.1007/s00248-020-01615-5. PMID 33021677. S2CID 222148939.
  85. ^ Engler A (1903). Syllabus der Pflanzenfamilien: eine Übersicht über das gesamte Pflanzensystem mit Berücksichtigung der Medicinal- und Nutzpflanzen nebst einer Übersicht über die Florenreiche und Florengebiete der Erde zum Gebrauch bei Vorlesungen und Studien über specielle und medicinisch-pharmaceutische Botanik (3rd ed.). Berlin: Borntraeger. doi:10.5962/bhl.title.22956.
  86. ^ Neuhauser S, Kirchmair M, Bulman S, et al. (23 February 2014). "Cross-kingdom host shifts of phytomyxid parasites". BMC Evolutionary Biology. 14 (33): 33. Bibcode:2014BMCEE..14...33N. doi:10.1186/1471-2148-14-33. PMC 4016497. PMID 24559266.
  87. ^ Sparrow FK (1960). Aquatic Phycomycetes. University of Michigan studies, Scientific series, v. 15. Ann Arbor, University of Michigan Press. doi:10.5962/bhl.title.5685. LCCN 59011269. OCLC 1362167.
  88. ^ Dick, Michael W. (2001). Straminipilous Fungi: Systematics of the Peronosporomycetes Including Accounts of the Marine Straminipilous Protists, the Plasmodiophorids and Similar Organisms (1 ed.). Springer Dordrecht. doi:10.1007/978-94-015-9733-3. ISBN 978-94-015-9733-3. S2CID 28755980.
  89. ^ Sierra R, Mauffrey F, Cruz J, Holzmann M, Gooday AJ, Maurer-Alcalá X, Thakur R, Greco M, Weiner AKM, Katz LA, Pawlowski J (2022). "Taxon-rich transcriptomics supports higher-level phylogeny and major evolutionary trends in Foraminifera" (PDF). Molecular Phylogenetics and Evolution. 174 (107546): 107546. Bibcode:2022MolPE.17407546S. doi:10.1016/j.ympev.2022.107546. PMID 35690380. S2CID 249608315.
  90. ^ Gooday AJ, Holzmann M, Schwarzgruber E, Cedhagen T, Pawlowski J (2022). "Morphological and molecular diversity of monothalamids (Rhizaria, Foraminifera), including two new species and a new genus, from SW Greenland". European Journal of Protistology. 86: 125932. doi:10.1016/j.ejop.2022.125932. PMID 36347189.
  91. ^ Sergeeva NG, Anikeeva OV (2021). "Vellaria solenta (Monothalamea: Allogromiidae) — new species of soft-walled foraminifera from Sivash Bay (the Sea of Azov)" (PDF). Invertebrate Zoology. 18 (2): 152–158. doi:10.15298/invertzool.18.2.06. S2CID 237368888.
  92. ^ Jump up to: a b c d e Siemensma F, Holzmann M, Apothéloz-Perret-Gentil L, Clauß S, Voelcker E, Bettighofer W, et al. (2021). "Broad sampling of monothalamids (Rhizaria, Foraminifera) gives further insight into diversity of non-marine Foraminifera". European Journal of Protistology. 77: 125744. doi:10.1016/j.ejop.2020.125744. PMID 33191053. S2CID 225144520.
  93. ^ Dellinger M, Labat A, Perrouault L, Grellier P (May 2014). "Haplomyxa saranae gen. nov. et sp. nov., a new naked freshwater foraminifer". Protist. 165 (3): 317–329. doi:10.1016/j.protis.2014.03.007. PMID 24810177.
  94. ^ Maldonado, Manuel; López-Acosta, María; Sitjà, Cèlia; Aguilar, Ricardo; García, Silvia; Vacelet, Jean (10 June 2013). "A giant foraminifer that converges to the feeding strategy of carnivorous sponges: Spiculosiphon oceana sp. nov. (Foraminifera, Astrorhizida)". Zootaxa. 3669 (4): 571–584. doi:10.11646/zootaxa.3669.4.9. hdl:10261/92975. PMID 26312358.
  95. ^ Jump up to: a b Nakamura Y, Tuji A, Kimoto K, Yamaguchi A, Hori RS, Suzuki N (2020). "Ecology, Morphology, Phylogeny and Taxonomic Revision of Giant Radiolarians, Orodaria ord. nov. (Radiolaria; Rhizaria; SAR)". Protist. 172 (3): 125808. doi:10.1016/j.protis.2021.125808. ISSN 1434-4610. PMID 34186475. S2CID 235686052.
  96. ^ Jump up to: a b Takahashi O, Maekawa T, Dumitrica P, Nguyen PD, Komatsu T (2022). "Latentifistularia and other radiolarian species from the lower Smithian (Lower Triassic) Lang Son Formation, NE Vietnam". Revue de micropaléontologie. 75. Bibcode:2022RvMic..7500610T. doi:10.1016/j.revmic.2022.100610. S2CID 246833665.
  97. ^ Noble P, Aitchison JC, Danelian T, Dumitrica P, Maletz J, Suzuki N, Cuvelier J, Caridroit M, O'Dogherty L (2017). "Taxonomy of Paleozoic radiolarian genera" (PDF). Geodiversitas. 39 (3): 419–502. doi:10.5252/g2017n3a4. S2CID 89647246.
  98. ^ Kawachi, Masanobu; Nakayama, Takuro; Kayama, Motoki; Nomura, Mami; Miyashita, Hideaki; Bojo, Othman; Rhodes, Lesley; Sym, Stuart; Pienaar, Richard N.; Probert, Ian; Inouye, Isao; Kamikawa, Ryoma (June 2021). "Rappemonads are haptophyte phytoplankton" (PDF). Current Biology. 31 (11): 2395–2403.e4. Bibcode:2021CBio...31E2395K. doi:10.1016/j.cub.2021.03.012. PMID 33773100. S2CID 232369943.
  99. ^ Zagumyonnyi DG, Radaykina LV, Tikhonenkov DV (2021). "Triangulopteris lacunata gen. et sp. nov. (Centroplasthelida), a New Centrohelid Heliozoan from Soil". Diversity. 13 (12): 658. doi:10.3390/d13120658.
  100. ^ Hausmann K, Weitere M, Wolf M, Arndt H (2002). "Meteora sporadica gen. nov. et sp. nov. (Protista incertae sedis) – an extraordinary free-living protist from the Mediterranean deep sea". European Journal of Protistology. 38 (2): 171–177. doi:10.1078/0932-4739-00872.
  101. ^ Jump up to: a b Shɨshkin, Yegor (2022). "Spironematella terricola comb. n. and Spironematella goodeyi comb. n. (Hemimastigida = Hemimastigea = Hemimastigophora) for Spironema terricola and Spironema goodeyi with diagnoses of the genus and family Spironematellidae amended". Zootaxa. 5128 (2): 295–297. doi:10.11646/zootaxa.5128.2.8. PMID 36101172. S2CID 252220401.
  102. ^ Heiss AA, Warring SD, Lukacs K, Favate J, Yang A, Gyaltshen Y, Filardi C, Simpson AG, Kim E (2021). "Description of Imasa heleensis, gen. nov., sp. nov. (Imasidae, fam. nov.), a Deep-Branching Marine Malawimonad and Possible Key Taxon in Understanding Early Eukaryotic Evolution". Journal of Eukaryotic Microbiology. 68 (2): e12837. doi:10.1111/jeu.12837. PMID 33274482. S2CID 227281992.
  103. ^ Stairs CW, Táborský P, Salomaki ED, Kolisko M, Pánek T, Eme L, Hradilová M, Vlček Č, Jerlström-Hultqvist J, Roger AJ, Čepička I (December 2021), "Anaeramoebae are a divergent lineage of eukaryotes that shed light on the transition from anaerobic mitochondria to hydrogenosomes", Current Biology, 31 (24): 5605–5612, Bibcode:2021CBio...31E5605S, doi:10.1016/j.cub.2021.10.010, PMID 34710348
  104. ^ Táborský, Petr; Pánek, Tomáš; Čepička, Ivan (2017). "Anaeramoebidae fam. nov., a Novel Lineage of Anaerobic Amoebae and Amoeboflagellates of Uncertain Phylogenetic Position". Protist. 168 (5): 495–526. doi:10.1016/j.protis.2017.07.005. PMID 28985627.
  105. ^ Yabuki A, Gyaltshen Y, Heiss AA, Fujikura K, Kim E (2018). "Ophirina amphinema n. gen., n. sp., a New Deeply Branching Discobid with Phylogenetic Affinity to Jakobids". Scientific Reports. 8 (1): 16219. Bibcode:2018NatSR...816219Y. doi:10.1038/s41598-018-34504-6. PMC 6212452. PMID 30385814.
  106. ^ Galindo LJ, Prokina K, Torruella G, López-García P, Moreira D (April 2023). "Maturases and Group II Introns in the Mitochondrial Genomes of the Deepest Jakobid Branch". Genome Biology and Evolution. 15 (4): evad058. doi:10.1093/gbe/evad058. PMC 10139444. PMID 37029959.
  107. ^ Pánek T, Táborský P, Pachiadaki MG, Hroudová M, Vlček Č, Edgcomb VP, Čepička I (2015). "Combined Culture-Based and Culture-Independent Approaches Provide Insights into Diversity of Jakobids, an Extremely Plesiomorphic Eukaryotic Lineage". Frontiers in Microbiology. 6: 1288. doi:10.3389/fmicb.2015.01288. PMC 4649034. PMID 26635756.
  108. ^ Strother, Paul K.; Taylor, Wilson A.; van de Schootbrugge, Bas; Leander, Brian S.; Wellman, Charles H. (2020). "Pellicle ultrastructure demonstrates that Moyeria is a fossil euglenid". Palynology. 44 (3): 461–471. doi:10.1080/01916122.2019.1625457.
  109. ^ Lax G, Keeling PJ (2023). "Molecular phylogenetics of sessile Dolium sedentarium, a petalomonad euglenid". The Journal of Eukaryotic Microbiology. 70 (e12991): e12991. doi:10.1111/jeu.12991. PMID 37424051.
  110. ^ Jump up to: a b c d e Lax G, Kolisko M, Eglit Y, Lee WJ, Yubuki N, Karnkowska A, Leander BS, Burger G, Keeling PJ, Simpson AG (2021). "Multigene phylogenetics of euglenids based on single-cell transcriptomics of diverse phagotrophs". Molecular Phylogenetics and Evolution. 159 (107088). Bibcode:2021MolPE.15907088L. doi:10.1016/j.ympev.2021.107088. PMID 33545276. S2CID 232047208.
  111. ^ Jump up to: a b Cavalier-Smith, Thomas; Chao, Ema E.; Vickerman, Keith (2016). "New phagotrophic euglenoid species (New genus Decastava; Scytomonas saepesedens; Entosiphon oblongum), Hsp90 introns, and putative euglenoid Hsp90 pre-mRNA insertional editing". European Journal of Protistology. 56: 147–170. doi:10.1016/J.EJOP.2016.08.002. PMID 27718409.
  112. ^ Jump up to: a b c d Lax G, Cho A, Keeling PJ (2023). "Phylogenomics of novel ploeotid taxa contribute to the backbone of the euglenid tree". The Journal of Eukaryotic Microbiology. 70 (e12973): e12973. doi:10.1111/jeu.12973. PMID 36912454.
  113. ^ Lax, Gordon; Lee, Won Je; Eglit, Yana; Simpson, Alastair (2019). "Ploeotids Represent Much of the Phylogenetic Diversity of Euglenids". Protist. 170 (2): 233–257. doi:10.1016/J.PROTIS.2019.03.001. PMID 31102975.
  114. ^ Jump up to: a b Cavalier-Smith, Thomas (2016). "Higher classification and phylogeny of Euglenozoa". European Journal of Protistology. 56: 250–276. doi:10.1016/J.EJOP.2016.09.003. PMID 27889663.
  115. ^ Lee WJ, Blackmore R, Patterson DJ (1999). "Australian records of two lesser known genera of heterotrophic euglenids – Chasmostoma Massart, 1920 and Jenningsia Schaeffer, 1918". Protistology. 1: 10–16.
  116. ^ Larsen, Jacob; Patterson, David J. (1990). "Some flagellates (Protista) from tropical marine sediments". Journal of Natural History. 24 (4): 801–937. Bibcode:1990JNatH..24..801L. doi:10.1080/00222939000770571.
  117. ^ Lax, Gordon; Simpson, Alastair G.B. (2020). "The Molecular Diversity of Phagotrophic Euglenids Examined Using Single-cell Methods". Protist. 171 (5). doi:10.1016/J.PROTIS.2020.125757. PMID 33126020.
  118. ^ Belyaev, Artem O.; Zagumyonnyi, Dmitriy G.; Mylnikov, Alexander P.; Tikhonenkov, Denis V. (2022). "The Morphology, Ultrastructure and Molecular Phylogeny of a New Soil-Dwelling Kinetoplastid Avlakibodo gracilis gen. et sp. nov. (Neobodonida; Kinetoplastea)". Protist. 173 (4): 125885. doi:10.1016/j.protis.2022.125885. PMID 35667307. S2CID 248711669.
  119. ^ Shishkin, Yegor; Drachko, Daria; Zlatogursky, Vasily V. (22 April 2021). "The smallest known heliozoans are the Erebor lineage (nom. clad. n.) inside Microheliella maris (Eukaryota, Diaphoretickes), with the amendation of M. maris diagnosis and description of Berkeleyaesol magnus gen. nov., comb. nov. (Eukaryota, incertae sedis)". International Journal of Systematic and Evolutionary Microbiology. 71 (4). doi:10.1099/ijsem.0.004776. PMID 33886450. S2CID 233370018.
  120. ^ Lee, Won Je (2002). "Some free-living heterotrophic flagellates from marine sediments of inchon and Ganghwa Island, Korea". Korean Journal of Biological Sciences. 6 (2): 125–143. doi:10.1080/12265071.2001.9647643.
  121. ^ Yamaguchi M, Mori Y, Kozuka Y, Okada H, Uematsu K, Tame A, Furukawa H, Maruyama T, Worman CO, Yokoyama K (2012). "Prokaryote or eukaryote? A unique microorganism from the deep sea". Journal of Electron Microscopy. 61 (6): 423–431. doi:10.1093/jmicro/dfs062. PMID 23024290.
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